Pollen D A, Ronner S F
Science. 1981 Jun 19;212(4501):1409-11. doi: 10.1126/science.7233231.
Adjacent simple cells recorded and "isolated" simultaneously from the same microelectrode placement were usually tuned to the same orientation and spatial frequency. The responses of the members of these "spatial frequency pairs" to drifting sine-wave gratings were cross-correlates. Within the middle range of the spatial frequency selectivity curves, the responses of the paired cells differed in phase by approximately 90 percent. This phase relationship suggests that adjacent simple cells tuned to the same spatial frequency and orientation represent paired sine and cosine filters in terms of their processing of afferent spatial inputs and truncated sine and cosine filters in terms of the output of simple cells.
从同一微电极放置位置同时记录并“分离”出的相邻简单细胞，通常会被调谐到相同的方向和空间频率。这些“空间频率对”的成员对漂移正弦波光栅的反应是相互关联的。在空间频率选择性曲线的中间范围内，配对细胞的反应在相位上相差约90%。这种相位关系表明，调谐到相同空间频率和方向的相邻简单细胞，就其对传入空间输入的处理而言，代表了配对的正弦和余弦滤波器；就简单细胞的输出而言，则代表了截断的正弦和余弦滤波器。
