Wood B
Department of Human Anatomy and Cell Biology, University of Liverpool, UK.
Bioessays. 1996 Dec;18(12):945-54. doi: 10.1002/bies.950181204.
The common ancestor of modern humans and the great apes is estimated to have lived between 5 and 8 Myrs ago, but the earliest evidence in the human, or hominid, fossil record is Ardipithecus ramidus, from a 4.5 Myr Ethiopian site. This genus was succeeded by Australopithecus, within which four species are presently recognised. All combine a relatively primitive postcranial skeleton, a dentition with expanded chewing teeth and a small brain. The most primitive species in our own genus, Homo habilis and Homo rudolfensis, are little advanced over the australopithecines and with hindsight their inclusion in Homo may not be appropriate. The first species to share a substantial number of features with later Homo is Homo ergaster, or 'early African Homo erectus', which appears in the fossil record around 2.0 Myr. Outside Africa, fossil hominids appear as Homo erectus-like hominids, in mainland Asia and in Indonesia close to 2 Myr ago; the earliest good evidence of 'archaic Homo' in Europe is dated at between 600-700 Kyr before the present. Anatomically modern human, or Homo sapiens, fossils are seen first in the fossil record in Africa around 150 Kyr ago. Taken together with molecular evidence on the extent of DNA variation, this suggests that the transition from 'archaic' to 'modern' Homo may have taken place in Africa.
现代人类与大猩猩的共同祖先据估计生活在500万至800万年前,但人类或原始人类化石记录中的最早证据是拉密达地猿,来自埃塞俄比亚一个450万年前的遗址。这个属之后是南方古猿属,目前已识别出其中的四个物种。它们都具有相对原始的颅后骨骼、带有扩大的咀嚼齿的齿列和较小的大脑。我们自己所属的人属中最原始的物种,能人及鲁道夫人,相比南方古猿进步不大,事后看来将它们归入人属可能并不合适。第一个与后来的人属有大量共同特征的物种是匠人,即“早期非洲直立人”,它大约在200万年前出现在化石记录中。在非洲以外,化石原始人类表现为类似直立人的原始人类,在亚洲大陆和印度尼西亚接近200万年前出现;欧洲“早期智人”的最早确凿证据可追溯到距今60万至70万年前。解剖学意义上的现代人,即智人,化石最早于约15万年前出现在非洲的化石记录中。结合关于DNA变异程度的分子证据,这表明从“早期智人”到“现代智人”的转变可能发生在非洲。