Zylberberg L
Histochem J. 1977 Jul;9(4):505-20. doi: 10.1007/BF01002979.
The lingual glands of amphibians are confined to the dorsal face of the tongue and are formed by invaginations of the lingual epithelium. The secretory products have a heterogeneous composition. Mucosubstances are usually associated with proteins. The secretory product of the outer glandular cells is rich in mucosubstances of an acidity varying with species. In Anourans and Urodeles, the secretory product of the tubes contains abundant proteins and, where present, mucosubstances are less abundant and less acid than those produced at the surface of the tongue. Proteins and mucosubstances coexist in the same secretory granules, which exhibit a more or less homogeneous appearance in Gymnophiona and in Anourans while having a complex structure in Urodeles. In spite of their primitive anatomical features, the amphibian lingual glands present histochemical signs of a complete cellular evolution.
两栖动物的舌腺局限于舌的背面，由舌上皮内陷形成。其分泌产物成分各异。黏液物质通常与蛋白质相关。外层腺细胞的分泌产物富含酸度因物种而异的黏液物质。在无尾目和有尾目中，导管的分泌产物含有丰富的蛋白质，且若存在黏液物质，其含量比舌表面产生的黏液物质少且酸性弱。蛋白质和黏液物质共存于相同的分泌颗粒中，这些颗粒在蚓螈目和无尾目中呈现出或多或少均匀的外观，而在有尾目中具有复杂的结构。尽管两栖动物舌腺具有原始的解剖特征，但它们呈现出细胞完全进化的组织化学迹象。
