Cohen M, Gerneke W H
J S Afr Vet Assoc. 1976 Mar;47(1):35-7.
An intermandibular glandular area, never before described in any other ungulate has been found in the steenbok (Raphicerus campestris). It consists of enlarged sebaceous and apocrine sweat glands situated in the upper two-thirds of the dermis. They produce a dirty white almost flaky odoriferous substance which clings to the hairs of the area and is easily rubbed off for marking territorial areas as well as for marking females during mating. The infraorbital gland produces a black secretion which is the combined secretion of melanaceous, branched alveolar, sebaceous and enlarged apocrine sweat glands. The function of this secretion is not known as it is not used for any obvious marking purposes.
在石羚（Raphicerus campestris）中发现了一个下颌间腺区，这在其他有蹄类动物中从未被描述过。它由位于真皮上三分之二处的增大的皮脂腺和顶泌汗腺组成。它们产生一种脏白色、几乎呈片状的有气味物质，这种物质附着在该区域的毛发上，很容易被擦掉，用于标记领地以及在交配时标记雌性。眶下腺产生一种黑色分泌物，它是黑色素、分支肺泡、皮脂腺和增大的顶泌汗腺的混合分泌物。这种分泌物的功能尚不清楚，因为它没有用于任何明显的标记目的。
