Knaff D B, Arnon D I
DEPARTMENT OF CELL PHYSIOLOGY, UNIVERSITY OF CALIFORNIA, BERKELEY.
Proc Natl Acad Sci U S A. 1969 Oct;64(2):715-22. doi: 10.1073/pnas.64.2.715.
It has generally been accepted that plant photosynthesis involves two light reactions, one that proceeds best in short-wavelength light and is identified with oxygen evolution (System II) and another that proceeds best in long-wavelength light and is identified with a cycle electron flow (System I). This paper presents a concept of three light reactions in photosynthesis, based on new evidence that System II comprises two rather than one short-wavelength light reaction. These appear to operate in series and to be connected by an electron transport chain peculiar to System II. Parallel to System II is the long-wavelength light reaction of System I.
一般认为,植物光合作用涉及两个光反应,一个在短波长光下进行得最好,与氧气释放有关(光系统II),另一个在长波长光下进行得最好,与循环电子流有关(光系统I)。本文基于新的证据提出了光合作用中三个光反应的概念,该证据表明光系统II包含两个而非一个短波长光反应。这两个反应似乎是串联运行的,并通过光系统II特有的电子传递链相连。与光系统II平行的是光系统I的长波长光反应。