Science. 1976 Apr 30;192(4238):461-3. doi: 10.1126/science.192.4238.461.
The number of decapod species associated with the coral Pocillopora damicornis is compared between two regions on the Pacific coast of Panama which are of the same geologic age but differ in environmental characteristics. The relationship between number of species and coral head size does not differ between the two regions but species composition among coral heads is more variable in the fluctuating environment. Thus there are more species (55 compared to 37) associated with corals in the fluctuating environment than in the constant environment. These data impugn the concept that environmental constancy increases species richness. They support the hypothesis that species equilibrium within habitats is maintained by measurable ecological factors-in this case, the effects of a natural physical disturbance.
与鹿角珊瑚（Pocillopora damicornis）相关的十足目物种数量在巴拿马太平洋沿岸的两个地区进行了比较，这两个地区具有相同的地质年龄，但环境特征不同。两个地区的物种数量与珊瑚头大小之间的关系没有差异，但在波动环境中，珊瑚头之间的物种组成更为多变。因此，在波动环境中与珊瑚相关的物种（55 种相比 37 种）更多。这些数据质疑了环境稳定性增加物种丰富度的概念。它们支持了这样一种假设，即在生境中物种的平衡是由可测量的生态因素维持的，在这种情况下，是自然物理干扰的影响。
