Botany Department, The Field Museum, 1400 S. Lake Shore Drive, Chicago, Illinois 60605-2496.
Mycologia. 2004 Mar-Apr;96(2):368-87.
The Sordariales is a taxonomically diverse group that has contained from seven to 14 families in recent years. The largest family is the Lasiosphaer-iaceae, which has contained between 33 and 53 genera, depending on the chosen classification. To determine the affinities and taxonomic placement of the Lasiosphaeriaceae and other families in the Sordariales, taxa representing every family in the Sordariales and most of the genera in the Lasiosphaeriaceae were targeted for phylogenetic analysis using partial sequences of the large-subunit (LSU) nrDNA. Based on molecular data, only genera within the families Chaetomiaceae, Lasiosphaeriaceae and Sordariaceae are retained within the redefined Sordariales. The order is a coherent group with morphologies that vary along well-defined lines, including large ascomata with large-celled membraneous or coriaceous walls and ascospores that show variation on a distinctive developmental theme, often with appendages or sheaths. The paraphyletic chaetomiaceous complex and the strongly supported Sordariaceae are nested among taxa traditionally placed in the Lasiosphaeriaceae. Analyses also indicate that 11 genera belong in the paraphyletic lasiosphaeriaceous complex. These taxa share a similar developmental pattern in their ascospore morphology that extends to the Sordariales as a whole. Based on these similarities in morphology, 13 additional genera are retained within the lasiosphaeriaceous complex and more than 35 genera have relationships in the order overall. Based on LSU data, 17 genera that have been assigned to the Lasiosphaeriaceae sensu lato are transferred to other families outside the Sordariales and 22 additional genera with differing morphologies subsequently are transferred out of the order. Two new orders, Coniochaetales and Chaetosphaeriales, are recognized for the families Coniochaetaceae and Chaetosphaeriaceae respectively. The Boliniaceae is accepted in the Boliniales, and the Nitschkiaceae is accepted in the Coronophorales. Annulatascaceae and Cephalothecaceae are placed in Sordariomycetidae inc. sed., and Batistiaceae is placed in the Euascomycetes inc. sed.
索兰盘菌目是一个分类多样的类群，近年来包含了 7 到 14 个科。最大的科是腔孢腔菌科，它包含了 33 到 53 个属，具体取决于所选的分类。为了确定腔孢腔菌科和索兰盘菌目中其他科的亲缘关系和分类位置，针对索兰盘菌目中的每个科以及腔孢腔菌科中的大多数属，使用 LSU nrDNA 的部分序列进行了系统发育分析。基于分子数据，只有 chaetomiaceae、lasiosphaeriaceae 和 sordariaceae 这三个科的属被保留在重新定义的索兰盘菌目中。该目是一个具有形态学一致性的类群，形态特征沿着明确的线变化，包括具有大细胞膜状或革质壁的大型子囊壳和具有独特发育主题的孢子，通常具有附属物或鞘。拟腔孢腔菌复杂类群和强烈支持的 sordariaceae 嵌套在传统上放置在腔孢腔菌科中的类群中。分析还表明，11 个属属于拟腔孢腔菌复杂类群。这些类群在其孢子形态上具有相似的发育模式，延伸到整个索兰盘菌目。基于这些形态学上的相似性，在腔孢腔菌复杂类群中保留了 13 个额外的属，在整个目中有 35 个以上的属具有关系。基于 LSU 数据，将被分配到广义腔孢腔菌科的 17 个属转移到索兰盘菌目以外的其他科，随后将 22 个具有不同形态的属转移出该目。分别为腔孢腔菌科和拟壳球腔菌科创建了两个新目，即Coniochaetales 和 Chaetosphaeriales。Boliniaceae 被接受为 Boliniales，Nitschkiaceae 被接受为 Coronophorales。Annulatascaceae 和 Cephalothecaceae 被归入 Sordariomycetidae inc. sed.，Batistiaceae 被归入 Euascomycetes inc. sed.。
