Faculté des Sciences, Université Mohamed V, Rabat, Morocco.
PLoS One. 2011;6(12):e26882. doi: 10.1371/journal.pone.0026882. Epub 2011 Dec 13.
New tetradactyl theropod footprints from Upper Jurassic (Oxfordian-Kimmeridgian) have been found in the Iouaridène syncline (Morocco). The tracksites are at several layers in the intermediate lacustrine unit of Iouaridène Formation. The footprints were named informally in previous works "Eutynichnium atlasipodus". We consider as nomen nudum.
METHODOLOGY/PRINCIPAL FINDINGS: Boutakioutichnium atlasicus ichnogen. et ichnosp. nov. is mainly characterized by the hallux impression. It is long, strong, directed medially or forward, with two digital pads and with the proximal part of the first pad in lateral position. More than 100 footprints in 15 trackways have been studied with these features. The footprints are large, 38-48 cm in length, and 26-31 cm in width.
CONCLUSIONS/SIGNIFICANCE: Boutakioutichnium mainly differs from other ichnotaxa with hallux impression in lacking metatarsal marks and in not being a very deep footprint. The distinct morphology of the hallux of the Boutakioutichnium trackmaker -i.e. size and hallux position- are unique in the dinosaur autopodial record to date.
在上侏罗统（牛津阶-卡米阶）的 Iouaridène 向斜（摩洛哥）发现了新的四足兽脚类足迹。足迹位于 Iouaridène 组中间湖泊单元的几个层位。在以前的研究中，这些足迹被非正式地命名为“Eutynichnium atlasipodus”。我们认为这是一个无记名的名字。
方法/主要发现：Boutakioutichnium atlasicus 化石属和种是新命名的，主要特征是大脚趾印痕。它很长，很强壮，指向内侧或前方，有两个趾垫，第一趾垫的近端部分位于外侧位置。有超过 100 个足迹在 15 条足迹中具有这些特征。这些足迹很大，长度为 38-48 厘米，宽度为 26-31 厘米。
结论/意义：Boutakioutichnium 主要与其他具有大脚趾印痕的足迹类型不同，它没有跖骨痕迹，并且不是很深的足迹。Boutakioutichnium 足迹制造者的大脚趾的明显形态——即大小和大脚趾位置——在迄今为止的恐龙后肢记录中是独一无二的。
