Kataev Boris M, Liang Hongbin
Zoological Institute, Russian Academy of Sciences, Universitetskaya nab. 1, St. Petersburg 199034, Russia.; Email:
Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China.; Email:
Zootaxa. 2015 Feb 19;3920(1):1-39. doi: 10.11646/zootaxa.3920.1.1.
A taxonomic review of 23 species of the subgenus Pseudoophonus Motschulsky, 1844, the genus Harpalus Latreille, 1802, occurring in China is given, and a key to these species is provided. The species are divided in three species groups and five subgroups, the distinctive characters of which are listed. The following new synonyms are established: Harpalus calceatus Duftschmid, 1812 = Anisodactylus propinquus Ballion, 1870, syn. n.; H. davidi (Tschitschérine, 1897) = H. kailiensis Huang, 1992, syn. n.; = H. adenticulatus Huang, 1992, syn. n.; = H. cilihumerus Huang, Hu & Sun, 1994, syn. n.; H. fokienensis Schauberger, 1930 = H. muciulus Huang, 1992, syn. n.; H. griseus (Panzer, 1796) = H. xinjiangensis Huang, Hu & Sun, 1994, syn. n.; H. hauserianus Schauberger, 1929 = H. disaogashimensis Huang, 1995, syn. n.; H. pastor pastor Motschulsky, 1844 = H. penglainus Huang, Hu & Sun, 1994, syn. n.; = H. chiloschizontus Huang, 1995, syn. n.; H. rufipes (DeGeer, 1774) = H. scabripectus Huang, Hu & Sun, 1994, syn. n.; H. singularis Tschitschérine, 1906 = H. chengjiangensis Huang, 1993, syn. n.; H. sinicus Hope, 1845 = H. periglabellus Huang, 1992, syn. n.; = H. longihornus Lei & Huang, 1997, syn. n.; and H. tridens Morawitz, 1862 = H. hypogeomysis Huang, 1993, syn. n.; = H. pilosus Huang, 1995, syn. n. Statuses of H. yinchuanensis Huang, 1993 and H. disimuciulus Huang, Lei, Yan & Hu, 1996 are discussed. Lectotypes are designated for H. capito Morawitz, 1862, H. japonicus Morawitz, 1862 and H. eous Tschitschérine, 1901. New data on distribution of Pseudoophonus species in China are provided. Harpalus babai Habu, 1973 is reported from China (Jiangxi) for the first time. The following taxa are recorded from the following Chinese provinces for the first time: H. ussuriensis Chaudoir, 1863 from Hunan; H. aenigma (Tschitschérine, 1897) from Hubei, Jiangxi, and Guangxi; H. pastor Motschulsky, 1844 from Beijing and Xizang; H. fokienensis Schauberger, 1930 from Anhui and Jiangxi; H. simplicidens Schauberger, 1929 from Beijing, Shanghai, and Fujian; H. coreanus (Tschitschérine, 1895) from Anhui, Henan, Zhejiang, Shanghai, Hubei, and Guangdong; H. suensoni Kataev, 1997 from Yunnan; H. sinicus Hope, 1845 from Hainan; H. davidi (Tschitschérine, 1897) from Zhejiang, Hunan, and Guizhou; H. pseudohauserianus Kataev, 2001 from Sichuan; H. indicus orientalis Kataev, 2014 from Xizang, Guangxi, and Guangdong; H. hauserianus Schauberger, 1929 from Xizang and Guizhou; H. singularis Tschitschérine, 1906 from Taiwan; and H. calceatus Duftschmid, 1812 from Guangxi. In addition, H. sinicus and H. singularis are reported from Laos for the first time.
对中国境内步甲属（Harpalus Latreille，1802）伪奥步甲亚属（Pseudoophonus Motschulsky，1844）的23个物种进行了分类学综述，并提供了这些物种的检索表。这些物种分为三个物种组和五个亚组，并列出了其显著特征。建立了以下新异名：Harpalus calceatus Duftschmid，1812 = Anisodactylus propinquus Ballion，1870，同义名；H. davidi (Tschitschérine，1897) = H. kailiensis Huang，1992，同义名； = H. adenticulatus Huang，1992，同义名； = H. cilihumerus Huang，Hu & Sun，1994，同义名；H. fokienensis Schauberger，1930 = H. muciulus Huang，1992，同义名；H. griseus (Panzer，1796) = H. xinjiangensis Huang，Hu & Sun，1994，同义名；H. hauserianus Schauberger，1929 = H. disaogashimensis Huang，1995，同义名；H. pastor pastor Motschulsky，1844 = H. penglainus Huang，Hu & Sun，1994，同义名； = H. chiloschizontus Huang，1995，同义名；H. rufipes (DeGeer，1774) = H. scabripectus Huang，Hu & Sun，1994，同义名；H. singularis Tschitschérine，1906 = H. chengjiangensis Huang，1993，同义名；H. sinicus Hope，1845 = H. periglabellus Huang，1992，同义名； = H. longihornus Lei & Huang，1997，同义名；以及H. tridens Morawitz，1862 = H. hypogeomysis Huang，1993，同义名； = H. pilosus Huang，1995，同义名。讨论了黄氏银川步甲（H. yinchuanensis Huang，1993）和黄氏双毛步甲（H. disimuciulus Huang，Lei，Yan & Hu，1996）的分类地位。为Harpalus capito Morawitz，1862、H. japonicus Morawitz，1862和H. eous Tschitschérine，1901指定了选模标本。提供了伪奥步甲在中国分布的新数据。首次报道巴氏步甲（Harpalus babai Habu，1973）在中国（江西）出现。首次从中国以下省份记录到以下分类单元：湘西步甲（H. ussuriensis Chaudoir，1863）来自湖南；谜步甲（H. aenigma (Tschitschérine，1897)）来自湖北、江西和广西；牧师步甲（H. pastor Motschulsky，1844）来自北京和西藏；福建步甲（H. fokienensis Schauberger，1930）来自安徽和江西；简单齿步甲（H. simplicidens Schauberger，1929）来自北京、上海和福建；朝鲜步甲（H. coreanus (Tschitschérine，1895)）来自安徽、河南、浙江、上海、湖北和广东；苏氏步甲（H. suensoni Kataev，1997）来自云南；中华步甲（H. sinicus Hope，1845）来自海南；戴氏步甲（H. davidi (Tschitschérine，1897)）来自浙江、湖南和贵州；拟豪氏步甲（H. pseudohauserianus Kataev，2001）来自四川；东方印度步甲（H. indicus orientalis Kataev，2014）来自西藏、广西和广东；豪氏步甲（H. hauserianus Schauberger，1929）来自西藏和贵州；奇异步甲（H. singularis Tschitschérine，1906）来自台湾；以及钙质步甲（H. calceatus Duftschmid，1812）来自广西。此外，首次报道中华步甲和奇异步甲在老挝出现。
