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来自中欧上贝里亚阶的石珊瑚及其与同期珊瑚组合的比较。

Scleractinian corals from the upper Berriasian of central Europe and comparison with contemporaneous coral assemblages.

作者信息

Baron-Szabo Rosemarie Christine

机构信息

Smithsonian Institution, Department of Invertebrate Zoology, MRC-163, Washington, DC, 20013, United States of America. Research Institute Senckenberg, Senckenberganlage 25, D-60325 Frankfurt/Main, Germany..

出版信息

Zootaxa. 2018 Feb 22;4383(1):1-98. doi: 10.11646/zootaxa.4383.1.1.

DOI:10.11646/zootaxa.4383.1.1
PMID:29689916
Abstract

Scleractinian coral faunas from the upper Berriasian part of the Oehrli Formation of western Austria (Vorarlberg) and eastern Switzerland (Canton of Appenzell) are taxonomically described for the first time. Furthermore, scleractinian corals of the upper Berriasian part of the Oehrli Formation of the Swiss Cantons of Nidwalden and Uri are revised based on the study of type material. Lectotypes are designated for the species Dimorphocoeniopsis alpina (Koby, 1896) and Pleurophyllia tobleri (Koby, 1896). Sixty-one species belonging to 43 genera and 18 families were identified, making the coral fauna of the upper Oehrli Formation by far the most diverse among the Berriasian assemblages: Actinastrea pseudominima (Koby), A. sp., Adelocoenia bulgarica (Toula)(new combination), A. hexaphyllia (d'Orbigny)(new combination), A. radisensis (d'Orbigny)(new combination), Allocoeniopsis luciensis (d'Orbigny), Amphiaulastrea sp., Astraeofungia cf. decipiens (Michelin), Cladophyllia conybearei Milne Edwards Haime, Columnocoenia ksiazkiewiczi Morycowa, Comoseris jireceki Toula, Complexastrea seriata Turnšek, C. lobata Geyer, Cycloria mariscali (Felix)(new combination), Dermosmilia capitata (Koby), D. simplex Koby, Dimorphastrea excavata d'Orbigny, D. explanata De Fromentel, Dimorphocoeniopsis alpina (Koby), Ellipsocoenia lorioli (Koby), Enallhelia compressa (Münster), E. rathieri d'Orbigny, Epistreptophyllum cf. densum Roniewicz, Fungiastraea moeschi (Koby), Heliocoenia corallina Koby, H. humberti Étallon, H. minima Sikharulidze, Heterocoenia cf. inflexa (Eichwald), Latiphyllia neocomiensis De Fromentel, Latiastrea mucronata Sikharulidze, Latomeandra sp., Meandrastrea rudis (De Fromentel)(new combination), M. cf. lamberti (Bataller), Meandrophyllia corrugata (Michelin), Microsolena major (Ferry), M. cf. subexcavata Eguchi, Mitrodendron cf. modicum Eliášová, Mixastraea polyseptata Morycowa, Montlivaltia arcuata Beauvais, M. kaufmanni Koby, M. truncata (Defrance, 1817), Myriophyllia cf. propria Sikharulidze, Paraclausastrea vorarlbergensis Baron-Szabo, Peplosmilia stutzi (Koby), Placocoenia heimi (Koby)(new combination), Placophyllia dianthus (Goldfuss), Plesiomontlivaltia paucisepta (Koby)(new combination), Pleurophyllia schmidti (Koby)(new combination), P. tobleri (Koby), Polyphylloseris icaunensis (d'Orbigny, 1850), Rhipidogyra cf. minima Koby, Stylangia cf. laddi Wells, Stylina pleionantha Meneghini, S. decipiens Étallon, S. cf. sparsa Trautschold, Stylosmilia alpina Koby, S. yabei Eguchi, Thecosmilia dichotoma (Koby), Th. sp., Trigerastraea gourdani (De Fromentel), and Vallimeandra cf. explanata (De Fromentel). Furthermore, five additional Berriasian coral faunas are reviewed and compared with the coral fauna of the upper Oehrli Formation. These five assemblages are from southern Ukraine (2 assemblages, consisting of five and 12 species, respectively), northern Tunisia (13 species), southern Spain (23 species), and central Tibet (11 species). Except for the faunas from central Tibet and northern Tunisia, the Berriasian coral assemblages are distinctly dominated by colonial species (70-95%); they represent largely isolated populations of mostly endemic species; and consist largely of genera that had already appeared in the Upper Jurassic (80-100%). On the species-level, however, with the exception of the coral assemblage of Spain, the Beriasian coral faunas are dominated by or completely consist of taxa that have their first occurrence in the Berriasian (54-100%). The Berriasian fauna of the upper Oehrli Formation described in this work contains nearly three times more species than found in the contemporaneous fauna of southern Spain which up to now was the largest known Berriasian fauna. Two of the upper Berriasian taxa described in the current work (Cycloria and Placocoenia) may be the first representatives of lineages that still occur today (Mussidae and Montastraeidae, respectively) as suggested by a recent study of the cox1 Intron in modern corals.

摘要

首次对奥地利西部(福拉尔贝格州)和瑞士东部(阿彭策尔州)厄尔利组上贝里亚阶的石珊瑚动物群进行了分类描述。此外,基于对模式标本的研究,对瑞士尼德瓦尔登州和乌里州厄尔利组上贝里亚阶的石珊瑚进行了修订。为Dimorphocoeniopsis alpina(科比,1896年)和Pleurophyllia tobleri(科比,1896年)这两个物种指定了选模。共鉴定出属于43个属、18个科的61个物种,这使得厄尔利组上部的珊瑚动物群成为贝里亚阶组合中迄今为止最为多样的:拟小放射星珊瑚(科比)、放射星珊瑚未定种、保加利亚异孔珊瑚(图拉)(新组合)、六叶异孔珊瑚(多尔比尼)(新组合)、拉迪异孔珊瑚(多尔比尼)(新组合)、卢西恩异孔珊瑚(多尔比尼)、双盘星珊瑚未定种、近似欺骗星珊瑚(米其林)、科尼比尔叶状珊瑚 米尔恩·爱德华兹和海姆、克西亚兹基柱孔珊瑚 莫雷科娃、伊雷采克多枝星珊瑚 图拉、细纹复星珊瑚 特尔恩舍克、叶状复星珊瑚 盖尔、马里斯卡利环星珊瑚(费利克斯)(新组合)、头状皮壳珊瑚(科比)、简单皮壳珊瑚 科比、挖掘双形星珊瑚 多尔比尼、扁平双形星珊瑚 德弗罗芒特尔、阿尔卑斯双形拟孔珊瑚(科比)、洛里奥利椭圆孔珊瑚(科比)、压缩全孔珊瑚(明斯特)、拉蒂埃全孔珊瑚 多尔比尼、近似致密表孔珊瑚 罗涅维茨、莫埃希真菌星珊瑚(科比)、珊瑚色太阳珊瑚 科比、洪贝蒂太阳珊瑚 埃塔隆、微小太阳珊瑚 西卡鲁利泽、近似弯曲异孔珊瑚(艾希瓦尔德)、新科米叶状珊瑚 德弗罗芒特尔、具短尖厚星珊瑚 西卡鲁利泽、宽叶珊瑚未定种、粗糙盘星珊瑚(德弗罗芒特尔)(新组合)、近似兰伯蒂盘星珊瑚(巴塔列尔)、波状曲叶珊瑚(米其林)、大型微管珊瑚(费里)、近似亚挖掘微管珊瑚 江口、近似小型线粒体树珊瑚 埃利亚绍娃、多隔混星珊瑚 莫雷科娃、弓形蒙利瓦珊瑚 博韦、考夫曼蒙利瓦珊瑚 科比、截形蒙利瓦珊瑚(德弗朗斯,1817年)、近似固有多叶珊瑚 西卡鲁利泽、福拉尔贝格副克劳斯塔珊瑚 巴伦 - 萨博、施图茨褶皮珊瑚(科比)、海姆板孔珊瑚(科比)(新组合)、石竹叶板叶珊瑚(戈德富斯)、少隔近蒙利瓦珊瑚(科比)(新组合)、施密德肋叶珊瑚(科比)(新组合)、托布勒肋叶珊瑚(科比)、伊坎多枝多叶星珊瑚(多尔比尼,1850年)、近似微小脊皱珊瑚 科比、近似拉迪斯廷珊瑚 韦尔斯、多花柱星珊瑚 梅内基尼、欺骗柱星珊瑚 埃塔隆、近似稀疏柱星珊瑚 特劳肖尔德、高山管星珊瑚 科比、矢部管星珊瑚 江口、二叉鞘孔珊瑚(科比)、鞘孔珊瑚未定种、古尔达尼三角星珊瑚(德弗罗芒特尔)、近似扁平瓦利珊瑚(德弗罗芒特尔)。此外,还对另外五个贝里亚阶珊瑚动物群进行了综述,并与厄尔利组上部的珊瑚动物群进行了比较。这五个组合分别来自乌克兰南部(两个组合,分别由5种和12种组成)、突尼斯北部(13种)、西班牙南部(23种)和西藏中部(11种)。除了来自西藏中部和突尼斯北部的动物群外,贝里亚阶珊瑚组合明显以群体物种为主(70 - 95%);它们主要代表了大多为特有物种的孤立种群;并且主要由上侏罗统已经出现的属组成(80 - 100%)。然而,在物种层面上,除了西班牙的珊瑚组合外,贝里亚阶珊瑚动物群以在贝里亚阶首次出现的分类单元为主或完全由这些分类单元组成(54 - 100%)。本文描述的厄尔利组上部的贝里亚阶动物群所含物种几乎是西班牙南部同期动物群的三倍,而西班牙南部动物群是迄今为止已知最大的贝里亚阶动物群。根据最近对现代珊瑚中细胞色素氧化酶亚基1内含子的研究,当前工作中描述的两个上贝里亚阶分类单元(环星珊瑚属和板孔珊瑚属)可能分别是现今仍存在的谱系(分别为蕈珊瑚科和蔷薇珊瑚科)的首个代表。

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