Manteuffel G, Kopp J, Himstedt W
Brain Behav Evol. 1986;28(4):186-97. doi: 10.1159/000118702.
Optokinetic nystagmus and after nystagmus were studied in six amphibian species, three urodeles and three anurans. It was demonstrated that two of the urodeles, Hydromantes italicus and Salamandra salamandra, display a relatively well-developed optokinetic after nystagmus, which is less pronounced in Bombina variegata and nearly absent in Bufo bufo, Rana temporaria, and Tylototriton verrucosus. These results indicate a certain degree of velocity storage in the optokinetic reflex of some amphibians.