Saha N, el Seikh F S
Am J Phys Anthropol. 1987 Jul;73(3):397-406. doi: 10.1002/ajpa.1330730310.
The distribution of ABO and Rhesus blood groups, serum haptoglobin, and transferrin; red cell glucose-6-phosphate dehydrogenase and acid phosphatase; and hemoglobin was studied among the two aboriginal negroid tribes (Nuba and Fur); the Nilotic tribe; five tribes of Arab ancestory; and a mixed group of other minor tribes of Arab origin. The Nilotic and Nuba tribes were genetically quite distinct from the rest, with lower R1, R2, and r in the Rhesus system and low HbS and Gd-. The Arab tribes had a genetic structure which was intermediate between that of the original negroid population of the Sudan and the Arabs to the north. However, some of the Arab tribes had special genetical characteristics, e.g., Messeria had high TfD1; both Messeria and Hawazma had high HbS and Gd-, while GdA was higher only in the Hawazma. The Gaalin had very low HbS, Ro, GdA, and Gd-, suggestive of less negroid admixture compared to Messeria and Hawazma. The Fur, though an aboriginal negroid tribe, had genetic characteristics similar to Arabs.
对两个原住民黑人部落（努巴族和富尔族）、尼罗特部落、五个阿拉伯裔部落以及一个由其他阿拉伯裔小部落组成的混合群体，研究了ABO和恒河猴血型、血清触珠蛋白和转铁蛋白；红细胞葡萄糖-6-磷酸脱氢酶和酸性磷酸酶；以及血红蛋白的分布情况。尼罗特部落和努巴部落在基因上与其他部落截然不同，在恒河猴血型系统中R1、R2和r比例较低，且血红蛋白S（HbS）和葡萄糖-6-磷酸脱氢酶（Gd-）水平较低。阿拉伯部落的基因结构介于苏丹原始黑人种群和北方阿拉伯人之间。然而，一些阿拉伯部落具有特殊的基因特征，例如，梅塞里亚部落转铁蛋白D1（TfD1）比例较高；梅塞里亚部落和哈瓦兹马部落的血红蛋白S和葡萄糖-6-磷酸脱氢酶比例都较高，而葡萄糖-6-磷酸脱氢酶A（GdA）仅在哈瓦兹马部落中比例更高。加林部落的血红蛋白S、Ro、葡萄糖-6-磷酸脱氢酶A和葡萄糖-6-磷酸脱氢酶比例非常低，这表明与梅塞里亚部落和哈瓦兹马部落相比，其黑人血统混合较少。富尔族虽然是一个原住民黑人部落，但其基因特征与阿拉伯人类似。
