Nabozhenko Maxim V, Bukejs Andris, Telnov Dmitry
Caspian Institute of Biological Resources of Dagestan Scientific Centre of the Russian Academy of Sciences, M. Gadzhiev str. 45, Makhachkala, Republic of Dagestan, 367000, Russia. Dagestan State University, 43a M. Gadzhiev Str., Makhachkala, Republic of Dagestan, 367000, Russia, [ORCID: 0000-0001-7914-7942].
Zootaxa. 2019 Mar 8;4565(2):zootaxa.4565.2.9. doi: 10.11646/zootaxa.4565.2.9.
Based on well preserved specimen, Gonialaena groehni gen. et sp. nov. (Coleoptera: Tenebrionidae) is described from Eocene Baltic amber and assigned to Gonialaenini trib. nov. within the subfamily Lagriinae due to the combination of the following characters: labrum elongate; antennal sensilla simple; procoxal cavities closed at least externally; narrow membranes present between abdominal ventrites 3-5; mesocoxal cavities partly closed by mesepimeron, trochantine present; ovipositor of primitive lagrioid type-digitate fourth coxite lobe with gonostyles apically. Gonialaena gen. nov. combines characters of various Lagriinae tribes: humeral angles rounded, formed by basally widened epipleura as in Goniaderini, Pycnocerini and Lupropini; narrow acute intercoxal process of abdominal ventrite 1 as in Belopini, Cossyphini and Lagriini; absence (or strong reduction) of metathoracic wings as in most Adeliini and some Lupropini; large, reniform eyes as in Goniaderini, Pycnocerini, Lagriini and many Adeliini; present intersegmental membranes between abdominal ventrites 3-5 as in all lagriine tribes except for Belopini and Cossyphini; subcylindrical terminal maxillary palpomere as in Belopini and Cossyphini. Thus, the position of Gonialaenini trib. nov. within Lagriinae is not fully understood, but combination of certain features (humeral angles rounded but distinct and vertical border present at elytral base, inconspicuous elytral striae, large reniform eyes, narrow intersegmental membranes) points on morphological similarity with Goniaderini, especially some Anaedus Blanchard, 1845 with indistinct elytral striae and related genera.
基于保存完好的标本，从始新世波罗的海琥珀中描述了新属新种格氏戈尼亚莱纳（鞘翅目：拟步甲科），并因其具有以下特征组合而被归入拟步甲亚科内的新族戈尼亚莱纳族：上唇细长；触角感器简单；前胸基节窝至少外部封闭；腹部腹板3 - 5之间有狭窄的膜；中胸基节窝部分被中后侧片封闭，有转节；产卵器为原始拟步甲类类型——第四腹板叶指状，顶端有生殖突。新属戈尼亚莱纳结合了不同拟步甲亚科族的特征：肩角圆润，由基部加宽的侧缘形成，如戈尼亚德族、扁角拟步甲族和卢普罗族；腹部腹板1的基节间突狭窄尖锐，如贝洛族、科斯西菲族和拟步甲族；后胸翅缺失（或强烈退化），如大多数阿德利族和一些卢普罗族；眼睛大，肾形，如戈尼亚德族、扁角拟步甲族、拟步甲族和许多阿德利族；腹部腹板3 - 5之间有节间膜，如除贝洛族和科斯西菲族外的所有拟步甲族；上颚末节圆柱形，如贝洛族和科斯西菲族。因此，戈尼亚莱纳族在拟步甲亚科内的位置尚未完全明确，但某些特征的组合（肩角圆润但明显，鞘翅基部有垂直边缘，鞘翅条纹不明显，眼睛大且肾形，节间膜狭窄）表明其在形态上与戈尼亚德族相似，尤其是一些1845年的布兰查德所描述的鞘翅条纹不明显的阿内杜斯及相关属。
