A reevaluation of the basal turtle from the Early-Middle Jurassic (Toarcian-Aalenian) of India, with descriptions of new material.

BACKGROUND

is an extinct turtle from the Early to Middle Jurassic Kota Formation of the Pranhita-Godavari Gondwana basin, India. The holotype and previously only known specimen is a partially eroded shell that had been collected near Kota village, north of Sironcha, in Maharashtra State. Phylogenetic analyses have consistently suggested placement at the base of the clade Mesochelydia.

METHODS

We here figure and describe the holotype of and two new specimens, which were collected from the Kota Formation near Kistapur village, Telengana State, about 60 km NW from the type locality. We furthermore explore the relationships of this fossil turtle by updating its scoring based on all available material in the most recent analysis of basal turtle relationships.

RESULTS

The revision of the holotype of provides minor adjustments to the morphology of this specimen, in particular recognition of a transverse break across the carapace, presence of only eight neurals, of which the eight is octagonal, and presence of a pathological element located between neurals VII and VIII. The new material provides new anatomical insights, in particular presence of a broad cervical, a vertebral V that inserts deeply into vertebral IV, narrow pleurals within increasingly short posteromedial contacts with the vertebrals towards the posterior, at least three pairs of musk duct foramina, and numerous insights into the morphology of the girdles and stylopodium. In combination, all material allows affirming the validity of with confidence. The phylogenetic analysis affirms the placement of as a basal mesochelydian, but cannot resolve its relationships relative to the roughly coeval and .