Schmidt Joachim, Will Kipling
University of Rostock, Institute of Biosciences, General and Systematic Zoology, Universitätsplatz 2, D-18055 Rostock, Germany..
Zootaxa. 2020 Jan 15;4722(4):zootaxa.4722.4.2. doi: 10.11646/zootaxa.4722.4.2.
The taxonomic position of Pristodactyla advena LeConte, 1848 (= "Acalathus" advena in the sense of previous authors) is discussed in light of a novel review of the morphological characters of Calathina and closely related taxa. All Acalathus Semenov, 1889 species and all other Dolichina share the distinct character state of having the sensory pits of the apical gonocoxites reduced or absent, while these pits are well-developed in P. advena. Therefore, P. advena is moved into the genus Calathus Bonelli, 1810 (Calathina), where it is placed in the molecular phylogeny of Ruiz et al. (2010). This move necessitates the description of Certocalathus subgen. n. for this species. Diagnostic characters of the new subgenus with respect to other Calathina and Dolichina beetles are presented. Given the lack of any clear synapomorphies, the subtribe Calathina is not obviously defined by morphology alone. However, recent molecular phylogenies indicate its monophyly is highly likely and therefore diagnostic morphological characters are polythetic. The states of selected morphological characters in Calathina and Dolichina taxa are provided as a summary for this and future studies.
根据对卡拉西纳属（Calathina）及近缘类群形态特征的全新综述，讨论了1848年勒孔特（LeConte）命名的普氏原伪叶甲（Pristodactyla advena）（= 之前作者所指的“棘伪叶甲属（Acalathus）”的普氏伪叶甲）的分类地位。1889年的所有棘伪叶甲属（Acalathus Semenov）物种及所有其他多丽伪叶甲属（Dolichina）物种都具有独特的特征状态，即顶端生殖基节的感觉窝减少或缺失，而在普氏原伪叶甲中这些感觉窝发育良好。因此，普氏原伪叶甲被移入1810年博内利（Bonelli）建立的卡拉西纳属（Calathus）（卡拉西纳亚属（Calathina）），并在鲁伊斯等人（Ruiz et al.，2010）的分子系统发育研究中得以定位。这一变动使得有必要为该物种描述新的亚属—— Certocalathus亚属。文中给出了该新亚属相对于其他卡拉西纳亚属和多丽伪叶甲属甲虫的鉴别特征。鉴于缺乏明显的共衍征，仅靠形态学无法明确界定卡拉西纳亚族。然而，近期的分子系统发育研究表明其单系性很可能成立，因此鉴别形态特征是多征的。文中提供了卡拉西纳亚属和多丽伪叶甲属类群中选定形态特征的状态，作为本研究及后续研究的总结。
