Sasakawa Kôji, Itô Hirotarô
Laboratory of Zoology, Department of Science Education, Faculty of Education, Chiba University, 1-33 Yayoicho, Inage-ku, Chiba-shi, Chiba, 263-8522 Japan.
1-14-16 Awayama, Niigata-shi, Niigata, 950-0843 Japan.
Zookeys. 2018 Mar 12(742):91-104. doi: 10.3897/zookeys.742.23752. eCollection 2018.
Bates, 1883, a Japanese endemic genus in the subtribe Synuchina (Coleoptera, Carabidae, Sphodrini), is revised taxonomically based mainly on the shape of the endophallus, a membranous inner sac everted from the aedeagus of the male genitalia. Three known species from central Honshu, Habu, 1954; Habu, 1978; and Habu, 1978, are re-defined based on this genital character, and five new species are described from the region: Sasakawa, . ., Sasakawa, . ., Sasakawa & Itô, . ., Sasakawa & Itô, . ., and Sasakawa & Itô, . . The observed interspecies differences in endophallus morphology are discussed in terms of the species-level phylogeny and genus-level taxonomy of .
Bates属，1883年，是日本特有的一个属，隶属于Synuchina亚族（鞘翅目，步甲科，Sphodrini族），主要基于阳茎内囊（从雄性生殖器的阳茎外翻出的膜质内囊）的形状进行了分类修订。根据这一生殖器特征，对本州中部已知的三个物种，即Habu，1954年；Habu，1978年；以及Habu，1978年，进行了重新定义，并从该地区描述了五个新物种：Sasakawa，……，Sasakawa，……，Sasakawa & Itô，……，Sasakawa & Itô，……，以及Sasakawa & Itô，……。从物种水平系统发育和该属的属级分类学角度讨论了观察到的阳茎内囊形态的种间差异。
