Perreault Ray T, Buckeridge John S
Jarreau Scientific, 6107 Prescott Road, Baton Rouge, Louisiana 70805, USA.
Zootaxa. 2019 Dec 18;4712(1):zootaxa.4712.1.2. doi: 10.11646/zootaxa.4712.1.2.
In North America, Paleogene Verrucidae are rarely encountered. Only a single named species has been previously discussed by Zullo Baum (1979), who provisionally assigned a Palaeocene form from North Carolina to Verruca rocana Steinmann, 1921. Eocene deposits in Washington State (USA) have now yielded two new species from intertidal to shallow water environments: Verruca gailgoedertae sp. nov. from the middle Eocene Crescent/McIntosh transition zone, and Verruca sorrellae sp. nov. from the upper Eocene to lowest Oligocene Gries Ranch and basal Lincoln Creek formations. Both species are characterized by punctate shell plates, and are placed in the lineage of Verruca stroemia (O.F. Müller, 1776). In Alabama, marls from the upper Yazoo Formation (Pachuta and Shubuta Members) have yielded Verruca alabamensis sp. nov., an uncommon deeper water form associated with abundant brachiopods and phosphate deposition. This last species shows no development of punctae and is not related to the lineage that leads directly to Verruca stroemia.
在北美，古近纪的藤壶科物种很少被发现。祖洛·鲍姆（1979年）之前仅讨论过一个已命名的物种，他将来自北卡罗来纳州的一种古新世形态临时归为1921年施泰因曼命名的罗氏藤壶（Verruca rocana）。美国华盛顿州的始新世沉积物现在已发现了两种来自潮间带到浅水环境的新物种：来自始新世中期新月/麦金托什过渡带的盖尔·戈德尔特氏藤壶（Verruca gailgoedertae）新种，以及来自始新世晚期至渐新世最底层的格里斯牧场和林肯溪组底部的索雷尔氏藤壶（Verruca sorrellae）新种。这两个物种的特征都是壳板有小孔，并被归入斯特罗米亚藤壶（Verruca stroemia，奥古斯特·弗里德里希·米勒，1776年）的谱系中。在阿拉巴马州，亚祖组上部（帕丘塔和舒布塔段）的泥灰岩产出了阿拉巴马藤壶（Verruca alabamensis）新种，这是一种罕见的深水物种，与丰富的腕足动物和磷酸盐沉积有关。最后这个物种没有小孔的发育，并且与直接通向斯特罗米亚藤壶的谱系无关。
