Evsyukov Aleksandr, Golovatch Sergei, Reip Hans, Vandenspiegel Didier
Don State Technical University, Department of Biology and General Pathology, Gagarin sq. 1, building 6, Rostov-on-Don 344003, Russia.
Zootaxa. 2020 May 14;4778(2):zootaxa.4778.2.2. doi: 10.11646/zootaxa.4778.2.2.
The generic classification of the Palaearctic tribe Leptoiulini is revised, with 11 genera being recognized. The main differences between the genera, all presented in a tabular form, lie in the structure of the 2nd and 7th leg-pairs of the male, coupled with gonopodal conformations: the presence/absence and the degree of development of flagella on the promeres, and of the phylacum and velum on the opisthomeres. Based on abundant new material, only three genera and seven species of Leptoiulini, all keyed, mapped and properly illustrated, are shown to occur in the Caucasus: Chatoleptophyllum flexum Golovatch, 1979, Kubaniulus gracilis Lohmander, 1936, K. lativelatus sp. nov., Leptoiulus hastatus Lohmander, 1932 (= L. disparatus Lohmander, 1936, syn. n.), L. tanymorphus (Attems, 1901), L. meskhii sp. nov., and L. gonopodialis sp. nov. All species are endemic or subendemic to the region, while Kubaniulus Lohmander, 1936 represents the only genus in the tribe which is restricted to the Caucasus, in particular, the western and central parts of the Caucasus Major. The distributions are mainly allopatric, but a few pairs of species are para- or even sympatric.
古北区瘦马陆族（Leptoiulini）的类属分类得到了修订，共确认了11个属。各属之间的主要差异均以表格形式呈现，体现在雄性第二和第七对足的结构以及生殖肢的形态上：前生殖肢上鞭毛的有无及发育程度，以及后生殖肢上叶状体和膜的情况。基于丰富的新材料，现已证明瘦马陆族仅有3个属和7个物种分布于高加索地区，文中对所有物种都进行了检索表编制、分布图绘制及详细配图说明：柔曲瘦叶马陆Chatoleptophyllum flexum Golovatch，1979；纤细库班马陆Kubaniulus gracilis Lohmander，1936；宽侧库班马陆K. lativelatus sp. nov.；戟形瘦马陆Leptoiulus hastatus Lohmander，1932（= L. disparatus Lohmander，1936，新异名）；异形瘦马陆L. tanymorphus（Attems，1901）；梅氏瘦马陆L. meskhii sp. nov.；以及具生殖肢瘦马陆L. gonopodialis sp. nov.。所有物种均为该地区特有或亚特有物种，而1936年的库班马陆属（Kubaniulus Lohmander）是该族中唯一局限于高加索地区的属，特别是大高加索山脉的西部和中部。这些物种的分布主要是异域分布，但也有少数物种对是邻域分布甚至同域分布。
