Zacca Thamara, Casagrande Mirna M, Mielke Olaf H H, Huertas Blanca, Espeland Marianne, Freitas AndrÉ V L, Willmott Keith R, Nakahara Shinichi, Lamas Gerardo
Departamento de Biologia Animal and Museu da Biodiversidade, Instituto de Biologia, Laboratório de Ecologia e Sistemática de Borboletas, Universidade Estadual de Campinas, Campinas, São Paulo, Brazil..
Zootaxa. 2020 Sep 29;4858(1):zootaxa.4858.1.1. doi: 10.11646/zootaxa.4858.1.1.
Vareuptychia Forster, 1964 stat. rest. is revalidated and comprises two species, V. similis (Butler, 1867) comb. rest. and V. themis (Butler, 1867) comb. nov. Vanima Zacca, Casagrande Mielke gen. nov. is described to contain Euptychia labe Butler, 1870 (the type species), E. palladia Butler, 1867 and E. lesbia Staudinger, [1886]. The taxonomy of these two genera was initially revised based on morphological and distributional data, and subsequently tested and supported with a Maximum Likelihood analysis using four genes (COI, GAPDH, RpS5 and EF1-a). Lectotypes are designated for Euptychia similis Butler, 1867, E. themis Butler, 1867, E. undina Butler, 1870 and E. lesbia Staudinger, [1886]. No DNA sequences were obtained for Euptychia cleophes Godman Salvin, 1889 but its transfer to Megisto Hübner, [1819] is supported by morphological evidence. For all taxa treated in this study, a taxonomic catalog, diagnosis, (re)description and illustrations of adults, venation and genitalia are provided, as well as comments on intraspecific variation, sexual dichromatism, ecology and distribution maps.
1964年福斯特恢复其有效地位的瓦雷普蒂奇亚属（Vareuptychia Forster, 1964 stat. rest.）现重新确认有效，包括两个物种，即V. similis（巴特勒，1867年）恢复组合和V. themis（巴特勒，1867年）新组合。新描述的瓦尼马属（Vanima Zacca, Casagrande Mielke gen. nov.）包含1870年的Euptychia labe巴特勒（模式种）、1867年的E. palladia巴特勒和[1886年]的E. lesbia施陶丁格。这两个属的分类最初基于形态学和分布数据进行修订，随后通过使用四个基因（COI、GAPDH、RpS5和EF1-a）的最大似然分析进行检验和支持。指定了1867年的Euptychia similis巴特勒、1867年的E. themis巴特勒、1870年的E. undina巴特勒和[1886年]的E. lesbia施陶丁格的选模标本。未获得1889年的Euptychia cleophes戈德曼·萨尔文的DNA序列，但形态学证据支持将其转移到[1819年]的Megisto许布纳属。对于本研究中处理的所有分类单元，提供了分类目录、诊断、成虫（重新）描述及成虫、脉序和生殖器的插图，以及关于种内变异、性二态性、生态学和分布图的注释。
