Ca de Massa, 25526 Llesp-El Pont de Suert, Lleida, Spain. Museu de CieÌncies Naturals (Zoologia), Passeig Picasso s/n, 08003 Barcelona, Spain..
Zootaxa. 2021 Feb 5;4926(2):zootaxa.4926.2.1. doi: 10.11646/zootaxa.4926.2.1.
We redefine the genus Troglocharinus Reitter, 1908 based on a phylogenetic analysis with a combination of mitochondrial and molecular data. We recovered the current Speonomites mengeli (Jeannel, 1910) and S. mercedesi (Zariquiey, 1922) as valid, separate species within the Troglocharinus clade, not directly related to Speonomites Jeannel, 1910, a finding corroborated by a detailed study of the male and female genitalia. In consequence, we reinstate Speonomus mercedesi Zariquiey, 1922 stat. nov. as a valid species, transfer both of them to the genus Troglocharinus, T. mengeli (Jeannel, 1910) comb. nov. and T. mercedesi (Zariquiey, 1922) comb. nov., and redescribe the genus. The study of new material from the distribution area of the former S. mengeli revealed the presence of two undescribed species, T. sendrai sp. nov. and T. fadriquei sp. nov., which we describe herein. We designate the lectotype of Speonomus vinyasi Escolà, 1971 to fix its identity, as among its syntypes there are two different species. In agreement with the results of the phylogenetic analyses we establish the synonymy between the genus Speonomites and Pallaresiella Fresneda, 1998 syn. nv.
我们基于线粒体和分子数据的系统发育分析，重新定义了 1908 年由 Reitter 建立的 Troglocharinus 属。我们恢复了现有的 Speonomites mengeli（Jeannel，1910）和 S. mercedesi（Zariquiey，1922）为有效、独立的物种，它们属于 Troglocharinus 分支，与 Speonomites Jeannel，1910 没有直接关系，这一发现得到了对雌雄生殖器详细研究的证实。因此，我们将 Speonomus mercedesi Zariquiey，1922 stat. nov. 恢复为有效物种，将它们都转移到 Troglocharinus 属中，即 T. mengeli（Jeannel，1910）comb. nov. 和 T. mercedesi（Zariquiey，1922）comb. nov.，并重新描述了该属。对来自前 S. mengeli 分布区的新材料的研究揭示了存在两个未描述的物种，T. sendrai sp. nov. 和 T. fadriquei sp. nov.，我们在此对它们进行了描述。我们指定 Speonomus vinyasi Escolà，1971 的模式标本来确定其身份，因为在其同型标本中有两个不同的物种。根据系统发育分析的结果，我们建立了 Speonomites 和 Pallaresiella Fresneda，1998 syn. nv. 的同义词关系。
