Harzhauser Mathias
Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria..
Zootaxa. 2021 Jan 5;4902(1):zootaxa.4902.1.1. doi: 10.11646/zootaxa.4902.1.1.
Evolutionary history, diversity and (paleo)geographic distribution of Cainozoic to present-day species of the Trochidae subfamilies Cantharidinae and Trochinae are discussed based on an extensive literature survey. In total, 393 species-level taxa, assigned to 24 genera and subgenera, are listed from the NE Atlantic, the E Atlantic, the North Sea, the (Proto)-Mediterranean Sea, the Central Paratethys Sea and the Eastern Paratethys Sea. Short diagnosis and subjective and objective junior synonyms for genus-level taxa are given. Stratigraphic ranges and geographic distribution are listed for species-level taxa. The European fossil record suggests a first major radiation during the middle Eocene and a second diversity pulse during the Miocene, when most extant genera were already present. At the species level, however, the present-day fauna is geologically very young, originating during the Pleistocene and Holocene. Overall, no convincing correlation of evolution and diversity of European Cantharidinae and Trochinae with major geodynamic events (e.g. Tethys Closure) can be observed. An exception is the somewhat overlooked spectacular radiation of Cantharidinae following the hydrological isolation of the Paratethys Sea during the late Miocene. The critical evaluation of the fossil record provides anchor points to test molecular phylogenies. A major discrepancy between both approaches appears only for Jujubinus, which suggests that Paleocene species have to be excluded from the genus. Gibbuliculus nov. gen. is introduced as new genus for a group of Oligocene to Pleistocene species, placed so far in "Colliculus" sensu auctores non Monterosato, 1888. Anceps siminescui nov. nom, Gibbuliculus saccoi nov. nom, Gibbula tavanii nov. nom., Gibbula s.l. lovellreevei nov. nom. and Gibbula s.l. steiningeri nov. nom. are proposed as new names for the preoccupied Trochus semistriatus Siminescu Barbu, 1940, Gibbula protumida Sacco, 1896, Gibbula minima Tavani, 1939, Trochus (Gibbula) reevei Harmer, 1923 and Trochus amedei bicincta Schaffer, 1912.
基于广泛的文献调查，讨论了海菊蛤亚科（Cantharidinae）和马蹄螺亚科（Trochinae）从新生代至今的物种演化历史、多样性及（古）地理分布。总共列出了来自东北大西洋、东大西洋、北海、（原）地中海、中副特提斯海和东副特提斯海的393个物种级分类单元，它们隶属于24个属和亚属。给出了属级分类单元的简短诊断以及主观和客观的次异名。列出了物种级分类单元的地层范围和地理分布。
欧洲的化石记录表明，在始新世中期出现了第一次主要的辐射，在中新世出现了第二次多样性脉冲，当时大多数现存属已经存在。然而，在物种层面，当今的动物群在地质上非常年轻，起源于更新世和全新世。总体而言，未观察到欧洲海菊蛤亚科和马蹄螺亚科的演化与多样性与主要地球动力学事件（如特提斯海关闭）之间有令人信服的相关性。一个例外是在中新世晚期副特提斯海水文隔离后，海菊蛤亚科出现了某种程度上被忽视的壮观辐射。对化石记录的批判性评估为检验分子系统发育提供了锚点。两种方法之间的一个主要差异仅出现在枣螺属（Jujubinus），这表明该属必须排除古新世的物种。
引入新属吉氏螺属（Gibbuliculus nov. gen.）来容纳一组渐新世至更新世的物种，这些物种迄今被置于“Colliculus”（非蒙特罗萨托，1888年作者意义上）。提出新名Anceps siminescui nov. nom、Gibbuliculus saccoi nov. nom、Gibbula tavanii nov. nom、Gibbula s.l. lovellreevei nov. nom和Gibbula s.l. steiningeri nov. nom，分别用于替代已被占用的Trochus semistriatus Siminescu Barbu, 1940、Gibbula protumida Sacco, 1896、Gibbula minima Tavani, 1939、Trochus (Gibbula) reevei Harmer, 1923和Trochus amedei bicincta Schaffer, 1912。
