Othenio Abel (1875-1946) and "the phylogeny of the parts".

The reconstruction of the evolutionary history of animal phyla was an integral part of Othenio Abel's paleobiology (paleozoology). Abel took issue with those phylogeneticists who, following the lead of Haeckel, would draw up phylogenetic trees on the basis of transformation series of singular characters considered to be of particular importance. Abel highlighted Louis Dollo's principle of the chevauchement des spécialisations (crossing of specializations), which transformed phylogenetics from a search for ancestor-descendant sequences to research into relative degrees of relationships. This replacement resolved the conflict, much discussed at the time, between the continuity of ancestor-descendant lineages and the discontinuity inherent in the natural (phylogenetic) system. Walter Zimmermann refined Abel's methodology, which he called character-phylogenetics (Merkmalsphylogenie), an approach that was eventually adopted by Willi Hennig.