Kaila Lauri
Zoological Museum, Finnish Museum of Natural History, University of Helsinki, Finland.
Cladistics. 2004 Aug;20(4):303-340. doi: 10.1111/j.1096-0031.2004.00027.x.
Phylogenetic relationships within the megadiverse lepidopteran superfamily Gelechioidea have been poorly understood and consequently the family level classification has been problematic. An analysis of phylogeny using 193 characters, including 241 informative character states, derived from larval, pupal and adult morphology and larval ecology, was performed to resolve the phylogeny of the Gelechioidea. 143 species representing the diversity of the putative Gelechioidea were included, supplemented with 13 species representing 11 other Ditrysian families. The monophyly of the Gelechioidea was supported, although only with homoplastic characters. The putative position of the Gelechioidea as the sister group of the Apoditrysia was not supported, since the Gelechioidea was nested within this clade. The Gelechioidea was divided into two main lineages: (1) the gelechiid lineage constituting Deoclonidae, Syringopainae, a re-composed Coleophoridae (including Coelopoetinae and Batrachedrinae as paraphyletic with Stathmopodinae, and Coleophorinae nested within it), Momphidae, Pterolonchidae, Scythrididae, Cosmopterigidae, and Gelechiidae, and (2) the oecophorid lineage constituting the "autostichid" family assemblage (including taxa formerly assigned to Autostichinae, Holcopogoninae, Symmocinae, Glyphidoceridae and Lecithoceridae), Xyloryctidae s.l. (including a paraphyletic Xyloryctidae of authors, some oecophorids of authors, Deuterogoniinae and Blastobasinae), Oecophoridae s.s., Amphisbatidae s.s., Carcinidae, Stenomati[n/d]ae, Chimabachidae and Elachistidae (including Depressariinae s.s., Telechrysis, Ethmiinae, Hypertrophinae s.l., miscellaneous "amphisbatids"sensu authors, Aeolanthinae, Parametriotinae, Agonoxeninae and Elachistinae). Detritivory/fungivory may have evolved only twice within Gelechioidea, though the evolution of larval food substrate use frequently reverses. To avoid an unnecessary further proliferation of names, it is recommended that no further family group names are introduced within the Gelechioidea, unless based on a rigorous analysis of inter-relationships.
在种类繁多的鳞翅目麦蛾总科内，系统发育关系一直未被充分理解，因此科级分类存在问题。利用193个特征（包括241个信息特征状态）进行系统发育分析，这些特征源自幼虫、蛹和成虫形态以及幼虫生态学，以解决麦蛾总科的系统发育问题。研究纳入了代表假定麦蛾总科多样性的143个物种，并补充了代表其他11个双孔亚目科的13个物种。麦蛾总科的单系性得到了支持，尽管只是通过同塑性特征。麦蛾总科作为无孔亚目的姐妹群的假定位置未得到支持，因为麦蛾总科嵌套在这个分支内。麦蛾总科分为两个主要谱系：（1）麦蛾谱系，包括澳麦蛾科、银麦蛾亚科、重新组合的鞘蛾科（包括巢蛾亚科和细蛾亚科，它们与纹蛾亚科构成并系群，且鞘蛾亚科嵌套其中）、麦蛾科、翼麦蛾科、潜蛾科、织蛾科和麦蛾科；（2）织蛾谱系，包括“异蛾科”科类组合（包括以前归入异蛾亚科、孔蛾亚科、圆翅异蛾亚科、谷蛾科和尖翅蛾科的类群）、广义木蛾科（包括作者所定义的并系性木蛾科、作者所定义的一些织蛾科类群、后孔蛾亚科和蛀果蛾亚科）、狭义织蛾科、双孔蛾科、蟹蛾科、窄蛾科、奇蛾科和巢蛾科（包括狭义凹翅巢蛾亚科、细蛾属、筛蛾亚科、广义肥须巢蛾亚科、作者所定义的各种“双孔蛾类”、风巢蛾亚科、副蛾亚科、异蛾亚科和巢蛾亚科）。在麦蛾总科内，食碎屑/食真菌习性可能仅进化了两次，尽管幼虫食物底物利用的进化经常发生逆转。为避免名称不必要的进一步增加，建议在麦蛾总科内不再引入更多科级类群名称，除非基于严格的亲缘关系分析。
