Tiburtini Manuel, Astuti Giovanni, Bartolucci Fabrizio, Casazza Gabriele, Varaldo Lucia, De Luca Daniele, Bottigliero Maria Vittoria, Bacchetta Gianluigi, Porceddu Marco, Domina Gianniantonio, Orsenigo Simone, Peruzzi Lorenzo
Department of Biology, University of Pisa, 56126 Pisa, Italy.
Botanic Garden and Museum, University of Pisa, 56126 Pisa, Italy.
Biology (Basel). 2022 Jul 14;11(7):1060. doi: 10.3390/biology11071060.
Three subspecies of are reported from Italy, two of which are considered endemic to the Apennines. The taxonomic value of these two taxa ( subsp. and subsp. ) is unclear. Moreover, the relationships between subsp. and Northern Italian populations-currently ascribed to subsp. -have never been addressed. Accordingly, we used an integrated taxonomic approach, including morphometry, seed morpho-colorimetry, karyology, molecular systematics (, , , , and region), and comparative niche analysis. According to our results, French-Northern Italian populations are clearly distinct from Apennine populations. In the first group, there is evidence which allows the recognition of s.str. (not occurring in Italy) and subsp. . In the second group, the two putative taxa endemic to the Northern Apennines cannot be separated, so a single subspecies is here recognized: subsp.
据报道，意大利有该物种的三个亚种，其中两个被认为是亚平宁山脉特有的。这两个分类单元（亚种 和 亚种 ）的分类学价值尚不清楚。此外，亚种 与目前归为 亚种 的意大利北部种群之间的关系从未得到探讨。因此，我们采用了综合分类学方法，包括形态测量学、种子形态比色法、核型分析、分子系统学（ 、 、 、 、 和 区域）以及比较生态位分析。根据我们的结果，法国 - 意大利北部种群与亚平宁种群明显不同。在第一组中，有证据表明可以识别出 指名亚种（在意大利未出现）和 亚种 。在第二组中，无法区分两个被认为是亚平宁山脉北部特有的分类单元，因此在此识别出一个单一的亚种： 亚种 。
