Richter J, Ullman S
Biol Cybern. 1986;54(4-5):313-7. doi: 10.1007/BF00318427.
Experiments by Schiller et al. have suggested that non-directional edge-specific simple cells are constructed from two directionally selective subunits with opposite preferred direction. This hierarchical notion was based on the fact that the responses of such units to edges moving in opposite directions are spatially displaced with respect to each other. An alternative explanation of the observed response separation is the delay between the responses of the center and surround mechanisms at the retinal level. Measurements of the response separation as a function of stimulus speed support this explanation and argues against the hierarchical notion of Schiller et al.
席勒等人的实验表明，非定向边缘特异性简单细胞是由两个具有相反偏好方向的方向选择性亚基构建而成。这一层次概念基于这样一个事实，即此类细胞对沿相反方向移动的边缘的反应在空间上相互错开。对观察到的反应分离的另一种解释是视网膜水平上中心和周边机制反应之间的延迟。作为刺激速度函数的反应分离测量结果支持了这一解释，并反驳了席勒等人的层次概念。
