Viraktamath C A, Webb M D, Yeshwanth H M
Department of Entomology; University of Agricultural Sciences; GKVK; Bengaluru 560065; India.
Department of Life Sciences (Insects); The Natural History Museum; Cromwell Road; London SW7 5BD; United Kingdom (Retired).
Zootaxa. 2025 Jan 10;5567(1):1-105. doi: 10.11646/zootaxa.5567.1.1.
A group of Asian Deltocephalinae leafhopper genera share one or more of the following features: head markings usually complex with a white diamond shaped marking apically, forewing with veins from outer apical cell (R1 and R2+3) not or variably strongly reflexed, subgenital plates moderately to greatly elongated to digitate apex, male pygofer with a dorsal or ventral process and aedeagus with one or two basal, lateral or ventral processes. This group, named here as the Monobazus group of genera, is treated in the present paper. The following genera are included: Banus Distant, Monobazus Distant, Osbornellus Ball, Mavromoustaca Dlabola, stat. nov., Phlogotettix Ribaut, Phlogothamnus Ishihara, Wanritettix Vilbaste and five new genera from the Indian subcontinent described in this paper. The new taxa recognized are: Banus curvatus sp. nov. (India: Karnataka, Kerala, Maharashtra, West Bengal), B. spiculatus sp. nov. (India: Karnataka, Kerala, Maharashtra, Tamil Nadu), Chandibanus gen. nov. with its type species Chandibanus dorsospinosus sp. nov. (India: Chandigarh), Cicurbanus gen. nov. (type species. Thamnotettix greeni Melichar), C. serratus sp. nov. (India: Karnataka, Kerala, Tamil Nadu), Lankabanus gen. nov. with its type species, Lankabanus attenuatus sp. nov., L. bispinosus sp. nov. (both from Sri Lanka), Madhyabus gen. nov. with its type species, M. reticulatus sp. nov. (India: Madhya Pradesh, Tamil Nadu), M. biligiris sp. nov. (India: Karnataka), Mavromoustaca brevispina sp. nov. (India: Punjab), M. channabasavanna sp. nov. (India: Karnataka), M. opressa sp. nov. (India: Himachal Pradesh, Mizoram, Uttarakhand), Monobazus angulatus sp. nov. (India: Karnataka, Uttarakhand), M. bannericus sp. nov. (India: Karnataka), M. biligiriensis sp. nov. (India: Karnataka, Tamil Nadu), M. bilineatus sp. nov. (India: Karnataka, Tamil Nadu), M. deccanensis sp. nov. (India: Andhra Pradesh, Karnataka, Kerala), M. longulus sp. nov. (India: Himachal Pradesh), M. nilgalaensis sp. nov. (Sri Lanka), M. paulus sp. nov. (India: Karnataka, Delhi), M. rotundus sp. nov. (Nepal), M. rubrostriatus sp. nov. (India: Karnataka, Kerala), M. sahyadricus sp. nov. (India: Karnataka, Kerala), M. shrunga sp. nov. (Nepal), M. sikkimensis sp. nov. (India: Sikkim), M. sudindicus sp. nov. (India: Tamil Nadu), Panghoella gen. nov. (type species, Banus maculosus Distant), P. nera sp. nov. (India: Himachal Pradesh, West Bengal. Myanmar. Nepal), P. sinuata sp. nov. (India: West Bengal) and Phlogothamnus umiamensis sp. nov. (India: Meghalaya). Osbornellus (Mavromoustaca) Dlabola is raised to genus level, Mavromoustaca Dlabola stat. nov. The following new combinations are proposed: Cicurbanus greeni (Melichar) comb. nov., C. cicur (Melichar) comb. nov. (both from Thamnotettix), C. notatus (Distant) comb. nov. (from Scaphoideus), Mavromoustaca arboropictus (Dlabola) comb. nov. (from Osbornellus), M. brunnescens (Pruthi) comb. nov. (from Thamnotettix), M. deviaticus (Dlabola) comb. nov. (from Osbornellus), M. macchiae (Lindberg) comb. nov. (from Circulifer), Monobazus aurantius (Xing & Li) comb. nov (from Osbornellus)., M. confuscus (Pruthi) comb. nov. (from Paralimnus), M. fanjingshanensis (Li) comb. nov. (from Phlogotettix), M. fuscovarius (Distant) comb. nov. (from Deltocephalus), M. fletcheri (Distant) comb. nov. (from Scaphoideus), M. circinatus (Yao, Zhang & Xing) comb. nov., M. acutaedeagus (Li) comb. nov., M. polymaculatus (Li & Song) comb. nov., M. productus (Li & Xing) comb. nov. and M. rugosus (Li) comb. nov. (all from Phlogothamnus), Panghoella conoidea (Xing & Li) comb. nov. (from Osbornellus), P. maculosa (Distant) comb. nov. (from Banus), P. suiyangensis (Xing & Li) comb. nov. (from Osbornellus), Phlogothamnus grimeus (Li & Wang) comb. nov. (from Phlogotettix). The following new synonyms are recognized: Thamnotettix cicur Melichar 1903 syn. nov. of T. greeni Melichar1903; Deltocephalus fuscovarius Distant 1918 syn. nov. of Monobazus dissimilis (Distant 1908). In addition, the Chinese genus Longicornus Li & Song is recorded from the Indian subcontinent for the first time (see Appendix) and its uncertain relationship to the Monobazus group is noted; two species from the Indian subcontinent are included: L. grandis (Pruthi) comb. nov. (from Monobazus) and L. longus Xing & Li. One species from China (Hainan Is.) is transferred to Monobazus, i.e., M. datianensis Xing, Dai & Li, 2008 (Scaphoidella) comb. nov. All taxa from the Indian subcontinent are diagnosed and/or described and illustrated and keys to genera and species are provided.
头部斑纹通常较为复杂，顶端有白色菱形斑纹；前翅从外端顶细胞发出的脉（R1和R2 + 3）不强烈或不同程度地强烈反折；亚生殖板适度到极大地延长，顶端呈指状；雄性尾节有背侧或腹侧突起，阳茎有一个或两个基部、侧面或腹侧突起。本文将这一组叶蝉属命名为莫诺巴祖斯属组，并对其进行了研究。其中包括以下属：巴努斯属（Distant）、莫诺巴祖斯属（Distant）、奥斯博内卢斯属（Ball）、马夫罗穆斯塔卡属（Dlabola，新地位）、焰叶蝉属（Ribaut）、焰条叶蝉属（Ishihara）、万日叶蝉属（Vilbaste）以及本文中描述的来自印度次大陆的五个新属。新确认的分类单元如下：弯巴努斯新种（印度：卡纳塔克邦、喀拉拉邦、马哈拉施特拉邦、西孟加拉邦）、刺巴努斯新种（印度：卡纳塔克邦、喀拉拉邦、马哈拉施特拉邦、泰米尔纳德邦）、钱迪巴努斯新属及其模式种背刺钱迪巴努斯新种（印度：昌迪加尔）、环斑巴努斯新属（模式种：绿氏拟菱纹叶蝉Melichar）、锯齿环斑巴努斯新种（印度：卡纳塔克邦、喀拉拉邦、泰米尔纳德邦）、兰卡巴努斯新属及其模式种细弱兰卡巴努斯新种、双刺兰卡巴努斯新种（均来自斯里兰卡）、中央巴努斯新属及其模式种网纹中央巴努斯新种（印度：中央邦、泰米尔纳德邦）、比利吉里斯中央巴努斯新种（印度：卡纳塔克邦）、短刺马夫罗穆斯塔卡新种（印度：旁遮普邦）、钱纳巴萨万纳马夫罗穆斯塔卡新种（印度：卡纳塔克邦）、压迫马夫罗穆斯塔卡新种（印度：喜马偕尔邦、米佐拉姆邦、北阿坎德邦）、角状莫诺巴祖斯新种（印度：卡纳塔克邦、北阿坎德邦）、班纳里库斯莫诺巴祖斯新种（印度：卡纳塔克邦）、比利吉里ensis莫诺巴祖斯新种（印度：卡纳塔克邦、泰米尔纳德邦）、双线莫诺巴祖斯新种（印度：卡纳塔克邦、泰米尔纳德邦）、德干莫诺巴祖斯新种（印度：安得拉邦、卡纳塔克邦、喀拉拉邦）、长形莫诺巴祖斯新种（印度：喜马偕尔邦）、尼尔加拉ensis莫诺巴祖斯新种（斯里兰卡）、保卢斯莫诺巴祖斯新种（印度：卡纳塔克邦、德里）、圆形莫诺巴祖斯新种（尼泊尔）、红纹莫诺巴祖斯新种（印度：卡纳塔克邦、喀拉拉邦）、西高止莫诺巴祖斯新种（印度：卡纳塔克邦、喀拉拉邦）、施伦加莫诺巴祖斯新种（尼泊尔）、锡金莫诺巴祖斯新种（印度：锡金）、南印度莫诺巴祖斯新种（印度：泰米尔纳德邦）、庞戈埃拉新属（模式种：斑巴努斯Distant）、内拉庞戈埃拉新种（印度：喜马偕尔邦、西孟加拉邦、缅甸、尼泊尔）、弯曲庞戈埃拉新种（印度：西孟加拉邦）以及乌米亚焰条叶蝉新种（印度：梅加拉亚邦）。奥斯博内卢斯（马夫罗穆斯塔卡）Dlabola被提升为属级，马夫罗穆斯塔卡Dlabola新地位。提出了以下新组合：绿氏环斑巴努斯（Melichar）新组合、西库尔环斑巴努斯（Melichar）新组合（均来自拟菱纹叶蝉属）、斑环斑巴努斯（Distant）新组合（来自舟形叶蝉属）、树栖马夫罗穆斯塔卡（Dlabola）新组合（来自奥斯博内卢斯属）、褐马夫罗穆斯塔卡（Pruthi）新组合（来自拟菱纹叶蝉属）、偏斜马夫罗穆斯塔卡（Dlabola）新组合（来自奥斯博内卢斯属）、马基亚马夫罗穆斯塔卡（Lindberg）新组合（来自环纹叶蝉属）、橙黄莫诺巴祖斯（Xing & Li）新组合（来自奥斯博内卢斯属）、混淆莫诺巴祖斯（Pruthi）新组合（来自副菱纹叶蝉属）、梵净山莫诺巴祖斯（Li）新组合（来自焰叶蝉属）、暗褐莫诺巴祖斯（Distant）新组合（来自小叶蝉属）、弗莱彻莫诺巴祖斯（Distant）新组合（来自舟形叶蝉属）、环状莫诺巴祖斯（Yao, Zhang & Xing）新组合、尖阳茎莫诺巴祖斯（Li）新组合、多斑莫诺巴祖斯（Li & Song）新组合、延长莫诺巴祖斯（Li & Xing）新组合以及皱纹莫诺巴祖斯（Li）新组合（均来自焰条叶蝉属）、锥形庞戈埃拉（Xing & Li）新组合（来自奥斯博内卢斯属）、斑纹庞戈埃拉（Distant）新组合（来自巴努斯属）、绥阳庞戈埃拉（Xing & Li）新组合（来自奥斯博内卢斯属）、粗糙焰条叶蝉（Li & Wang）新组合（来自焰叶蝉属）。确认了以下新异名：1903年的西库尔拟菱纹叶蝉Melichar是1903年绿氏拟菱纹叶蝉Melichar的新异名；1918年的暗褐小叶蝉Distant是2008年不同莫诺巴祖斯（Distant）新异名。此外，中国的长角叶蝉属Li & Song首次在印度次大陆被记录（见附录），并指出其与莫诺巴祖斯属组的关系不确定；包括来自印度次大陆的两个物种：大长角叶蝉（Pruthi）新组合（来自莫诺巴祖斯属）和长形长角叶蝉Xing & Li。来自中国（海南）的一个物种被转移到莫诺巴祖斯属，即2008年的大田莫诺巴祖斯Xing, Dai & Li（舟形叶蝉属）新组合。对来自印度次大陆的所有分类单元进行了诊断和/或描述及绘图，并提供了属和种的检索表。
