Shamshev Igor V, Vasilenko Dmitry V, Perkovsky Evgeny E
Zoological Institute; Russian Academy of Sciences; Universitetskaya nab. 1; St. Petersburg 199034; Russia..
Cherepovets State University; Lunacharsky Prospect 5; Cherepovets; Vologda Region; 162600; Russia.; Borissiak Paleontological Institute; Russian Academy of Sciences; Profsoyuznaya 123; Moscow; 117647; Russia..
Zootaxa. 2025 Jan 23;5575(2):283-294. doi: 10.11646/zootaxa.5575.2.4.
The family Hybotidae (Insecta: Diptera, Empidoidea) is recorded for the first time from the bio-inclusions of Sakhalinian amber (Russia). Almost all inclusions studied belong to the subfamily Tachydromiinae. First results suggest that tachydromiines were not uncommon in a palaeo-biotope, where a resin known nowadays as Sakhalinian amber was produced. A new genus and species, Eoplatypalpus sachalinensis gen. et sp. nov., are described and illustrated. In addition, the inclusions of two tachydromiine species of unclear genera (or genus) have been examined but not named. Preliminarily, the palaeo-fauna of Tachydromiinae of Sakhalinian amber is characterised by the presence of only primitive representatives. They show an affinity with the most primitive extant genera of the subfamily (such as Symballophthalmus Becker and Megagrapha Melander) and with the group of extinct genera known from Upper Cretaceous inclusions (Cretoplatypalpus group of genera). The paleontological records of Tachydromiinae are briefly discussed. In addition, it is shown that Symballophthalmus clavilabrosus Hong described from Ypresian Fushun amber of China actually belongs to a different, as of yet uncertain genus. Consequently, fossils of the primitive extant genus Symballophthalmus remain unknown. An identification key to fossil genera of Tachydromiinae (exclusive of Drapetidini) is provided.
巨蚊科（昆虫纲：双翅目，舞虻总科）首次在萨哈林琥珀（俄罗斯）的生物包体中被记录。几乎所有研究的包体都属于长足虻亚科。初步结果表明，长足虻在一个古生物群落中并不罕见，如今被称为萨哈林琥珀的树脂就是在那里产生的。描述并图示了一个新属新种，即萨哈林始宽须舞虻 Eoplatypalpus sachalinensis gen. et sp. nov.。此外，还检查了两个属（或类属）不明的长足虻物种的包体，但未命名。初步来看，萨哈林琥珀中长足虻亚科的古动物群仅以原始代表物种为特征。它们与该亚科现存最原始的属（如 Becker 的 Symballophthalmus 和 Melander 的 Megagrapha）以及从白垩纪晚期包体中已知的灭绝属类群（Cretoplatypalpus 属类群）有亲缘关系。简要讨论了长足虻亚科的古生物学记录。此外，研究表明，从中国始新世抚顺琥珀中描述的 Hong 的具棒唇 Symballophthalmus clavilabrosus 实际上属于一个不同的、尚未确定的属。因此，现存原始属 Symballophthalmus 的化石仍然未知。提供了一份长足虻亚科（不包括 Drapetidini）化石属的鉴定检索表。
