Kaplan N L, Brookfield J F
Theor Popul Biol. 1983 Jun;23(3):273-80. doi: 10.1016/0040-5809(83)90018-7.
The behavior of the frequency spectrum of sites of transposable elements for the model proposed by Langley, Brookfield, and Kaplan ((1983) Genetics 104) is studied when some of the sites have weak additive selection against the host. In particular, it is shown that under certain conditions, the expected homozygosity at equilibrium will always be larger when there is variation in selection between sites than when such variation is absent.