Bakulina E D, Sidorova N V
Ontogenez. 1976;7(4):368-72.
The low percentage of larval hatching both in the initial lines and the hybrids of direct and back crosses was shown to be due to the low fertilizability of eggs. A study of early embryogenesis, up to the gastrulation, has shown that the F1 hybrids (female D. virilis x male D. littoralis) develop at a lesser rate than both the parental species. Some embryos of these latter attain the stage of blastoderm syncytium for 2 hrs whereas the hybrid embryos attain only the stage of polynuclear syncytium.
研究表明，初始品系以及正反交杂种中低比例的幼虫孵化是由于卵的低受精能力所致。对直至原肠胚形成阶段的早期胚胎发育的研究表明，F1杂种（雌性粗壮果蝇×雄性海滨果蝇）的发育速度比双亲物种都要慢。双亲物种的一些胚胎在2小时内就能达到囊胚层合胞体阶段，而杂种胚胎仅能达到多核合胞体阶段。
