Pavlinov I Ia
Zh Obshch Biol. 2007 Jan-Feb;68(1):19-34.
Any research activity is conducted within the framework of a cognitive situation which is defined by certain basic assumptions about ontology of the portion of the objective world under investigation. From the standpoint of the non-classical scientific epistemology, a part of that situation is constituted by personal knowledge which is formed by a set of thinking (cognitive) styles. The scholastic thinking existing in taxonomy and phylogenetics is considered as an example showing unavoidability of such styles in the natural history knowledge. It is initially rooted in the antic, mythological by its essence, persuasion of isomorphism between movements of the objective reality and of the mind. The instrumentalism entailed by scholastic thinking is based on the mythologeme according to which the "right method" par excellence can lead to the "right knowledge". That is why any disputes between different numerical methods of phylogenetic reconstructions are vain: their validity could be assessed not formally but within particular cognitive situations formed by particular basic models of the phylogenesis. Phylogenetic thinking is of the key importance in evolutionary biology and has great impact on various fields of biology based on it. It is pretty mythological because of non-observability of the phylogenesis: the latter is rather "thinked-in" in the objective world then is induced from the observed facts. It constitutes a part of the evolutionary thinking considering mainly macroevolutionary trends and stressing the initial causes in the structure of causal relations in the analyses of the diversity of organisms. The "tree thinking" of O'Hara is its rough operational equivalent. Relation between phylogenetic thinking and some other styles are considered, which are population, phenetic, typological, and epigenetic ("developmental" of O'Hara). Phylogenetic thinking makes it obliged inclusion of the initial causes in the explanatory models which underlie adaptive and functional peculiarities of organisms, as well as of the entire structure of the biodiversity. It manifests itself in such kind of models through uncovering the phylogenetic signal. This thinking style has great effect on understanding of the ontology of taxa and acknowledges the objective status of the phylogenetic pattern. It is intrinsically included in the argumentation schemes of constructional morphology, comparative phylogenetics. The central metaphor of the phylogenetics is the Tree of Life. Emagination of its unity and uniquiness is of naturphilosophical nature. From the contemporary epistemological standpoint, it should be considered as a generalization upon partial hypotheses of evolution of particular structures each corresponding to certain consideration aspect of the global phylogenesis. Acknowledging of multi-aspectness of the phylogenesis constitutes one of the important points of modern phylogenetic thinking. As different semogeneses are incompletely congruent, the Tree of Life is less certain than each of the initial hypotheses. Any attempt to make it more resolved would lead to its reduction to any of the particular semogenetic scheme (i.e. to a "gene tree") or to its "decay" into several trees each corresponding to a particular consideration aspect of the global Tree.
任何研究活动都是在一种认知情境的框架内进行的,这种认知情境是由关于所研究的客观世界部分的本体论的某些基本假设所定义的。从非经典科学认识论的角度来看,这种情境的一部分是由个人知识构成的,而个人知识是由一组思维(认知)方式形成的。分类学和系统发育学中存在的经院式思维被视为一个例子,表明这种思维方式在自然历史知识中是不可避免的。它本质上最初植根于古代神话,即相信客观现实的运动与思维的同构性。经院式思维所蕴含的工具主义基于这样一个神话主题,即卓越的“正确方法”能够带来“正确知识”。这就是为什么系统发育重建的不同数值方法之间的任何争论都是徒劳的:它们的有效性不能从形式上评估,而只能在由特定系统发育基本模型形成的特定认知情境中进行评估。系统发育思维在进化生物学中至关重要,并对基于此的生物学各个领域产生重大影响。由于系统发育不可观测,它相当具有神话色彩:后者更多是在客观世界中“被构想出来”,而不是从观察到的事实中推导出来的。它构成了进化思维的一部分,主要考虑宏观进化趋势,并在分析生物多样性时强调因果关系结构中的初始原因。奥哈拉的“树形思维”是其大致的操作等效物。文中考虑了系统发育思维与其他一些思维方式的关系,这些思维方式包括群体思维、表型思维、类型学思维和表观遗传学思维(奥哈拉的“发育”思维)。系统发育思维使得在解释模型中必须纳入初始原因,这些解释模型是生物体适应性和功能特性以及生物多样性整体结构的基础。它通过揭示系统发育信号在这类模型中得以体现。这种思维方式对理解分类单元的本体论有很大影响,并承认系统发育模式的客观地位。它本质上包含在建构形态学、比较系统发育学的论证方案中。系统发育学的核心隐喻是生命之树。对其统一性和独特性的想象具有自然哲学的性质。从当代认识论的角度来看,它应被视为对特定结构进化的部分假设的概括,每个假设对应于全球系统发育的特定考虑方面。承认系统发育的多面性是现代系统发育思维的重要要点之一。由于不同的生源说并不完全一致,生命之树比任何一个初始假设都更不确定。任何使其更清晰的尝试都会导致它简化为任何一个特定的生源说方案(即“基因树”),或者“分解”为几棵树,每棵树对应于全球生命之树的一个特定考虑方面。