Pape H
Institut für Geophysik, Clausthal-Zellerfeld, FRG.
Genetica. 1990;80(2):115-28. doi: 10.1007/BF00127132.
The black and white spotting patterns of Landseer dogs are divided into qualitatively recognizable phenotypic classes. Breeding data were obtained from the Swiss Dog Stud Book (SHSB) and from breeders' recent records. A plausible interpretation assumed qualitative inheritance of the generally accepted piebald spotting gene sp1 with at least two modifiers, s2 and s3. The modifier genes are regarded as minor spotting genes and may be responsible for white markings in the related Newfoundland breed which has been cross-bred with Landseers. The proposed scheme of polygenic inheritance can also be applied to the piebald spotting pattern of Holstein-Friesian cattle, using breeding data from literature.
兰西尔犬的黑白斑点模式可分为可定性识别的表型类别。繁殖数据来自瑞士犬类育种登记簿（SHSB）以及育种者的近期记录。一种合理的解释假定普遍认可的花斑斑点基因sp1具有至少两个修饰基因s2和s3的定性遗传。修饰基因被视为次要斑点基因，可能与已与兰西尔犬杂交的相关纽芬兰犬品种中的白色斑纹有关。利用文献中的育种数据，所提出的多基因遗传方案也可应用于荷斯坦 - 弗里生奶牛的花斑斑点模式。
