Grismer L Lee, Wood Perry L Jr, Thura Myint Kyaw, Quah Evan S H, Grismer Marta S, Murdoch Matthew L, Espinoza Robert E, Lin Aung
Herpetology Laboratory, Department of Biology, La Sierra University, 4500 Riverwalk Parkway, Riverside, California 92515, USA..
Zootaxa. 2018 Jul 19;4446(4):477-500. doi: 10.11646/zootaxa.4446.4.4.
A phylogenetic taxonomic analysis indicates that a newly discovered population of Cyrtodactylus from the vicinity of Ywangan Town in the Shan Hills, Shan State, Myanmar is a new species (C. ywanganensis sp. nov.) and the earliest diverging member of the linnwayensis group within the previously defined Indochinese clade. The DIVALIKE+J model of a BioGeoBEARS biogeographic analysis indicates that the Indochinese clade evolved in the Shan Hills and Salween Basin of eastern Myanmar and dispersed into Indochina on at least three separate occasions from 18.6-13.4 mya. Once there, uplift of the Tenasserim Mountains and Thai Highlands created the intermedius group, the oldhami group, and C. tigroides of western and southern Thailand which form sister lineages to the linnwayensis group, yathepyanensis group, and the sinyineensis group, respectively, of eastern Myanmar. Diverging lineages within the Indochinese clade highlight the importance of the Thai Highlands and Tenasserim Mountains in that group's evolution and speciation. The discovery of C. ywanganensis sp. nov. in karstic habitats in the Shan Hills continues to underscore the unrealized karst-associated herpetological diversity of this vast, relatively unexplored, upland region and the need for additional field studies.
系统发育分类分析表明，在缅甸掸邦掸邦山区Ywangan镇附近新发现的弯脚虎种群是一个新物种（Ywangan弯脚虎新种），也是先前定义的印度支那分支中linnwayensis组最早分化的成员。BioGeoBEARS生物地理分析的DIVALIKE+J模型表明，印度支那分支在缅甸东部的掸邦山区和萨尔温江盆地进化，并在1860 - 1340万年前至少三次分别扩散到印度支那。到达那里后，丹那沙林山脉和泰国高地的隆升形成了中间组、奥氏组以及泰国西部和南部的虎纹弯脚虎，它们分别是缅甸东部linnwayensis组、yatethepyanensis组和sinyineensis组的姐妹谱系。印度支那分支内不同的谱系突出了泰国高地和丹那沙林山脉在该组进化和物种形成中的重要性。在掸邦山区岩溶栖息地发现Ywangan弯脚虎新种，继续凸显了这片广阔、相对未被探索的高地地区与岩溶相关的未被认识的爬行动物多样性，以及进行更多实地研究的必要性。
