Reichardt W E, Schlögl R W
Max-Planck-Institut für biologische Kybernetik, Tübingen, Federal Republic of Germany.
Biol Cybern. 1988;60(1):23-35. doi: 10.1007/BF00205969.
The local extraction of motion information from brightness patterns by individual movement detectors of the correlation-type is considered in the first part of the paper. A two-dimensional field theory of movement detection is developed by treating the distance between two adjacent photoreceptors as a differential. In the first approximation of the theory we only consider linear terms of the time interval between the reception of a contrast element and its delayed representation by the detector and linear terms of the spatial distances between adjacent photoreceptors. As a result we may neglect terms of higher order than quadratic in a Taylor series development of the brightness pattern. The responses of pairs of individual movement detectors are combined to a local response vector. In the first approximation of the detector field theory the response vector is proportional to the instantaneous pattern velocity vector and linearly dependent on local properties of the moving pattern. The linear dependence on pattern properties is represented by a two by two tensor consisting of elements which are nonlinear, local functionals of the moving pattern. Some of the properties of the tensor elements are treated in detail. So, for instance, it is shown that the off-diagonal elements of the tensor disappear when the moving pattern consists of x- and y-dependent separable components. In the second part of the paper the tensor relation leading to the output of a movement detector pair is spatially integrated. The result of the integration is an approximation to a summation of the outputs of an array of detector pairs. The spatially integrated detector tensor relates the translatory motion vector to the resultant output vector. It is shown that the angle between the motion vector and the resultant output vector is always smaller than +/- 90 degrees whereas the angle between the motion vector and local response vectors, elicited by detector pairs, may cover the entire angular range. In the discussion of the paper the limits of the field theory for motion computation as well as its higher approximations are pointed out in some detail. In a special chapter the dependence of the detector response on the pattern properties is treated and in another chapter questions connected with the so called aperture problem are discussed. Furthermore, properties for compensation of the pattern dependent deviation angle by spatial physiological integration are mentioned in the discussion.
本文第一部分探讨了相关型个体运动探测器从亮度模式中局部提取运动信息的问题。通过将两个相邻光感受器之间的距离视为微分,建立了运动检测的二维场论。在该理论的一阶近似中,我们仅考虑对比度元素接收与其在探测器中延迟表示之间的时间间隔的线性项,以及相邻光感受器之间空间距离的线性项。因此,在亮度模式的泰勒级数展开中,我们可以忽略高于二次的项。将成对个体运动探测器的响应组合成一个局部响应向量。在探测器场论的一阶近似中,响应向量与瞬时模式速度向量成正比,并且线性依赖于运动模式的局部属性。对模式属性的线性依赖由一个二阶张量表示,该张量的元素是运动模式的非线性局部泛函。详细讨论了张量元素的一些属性。例如,当运动模式由与x和y相关的可分离分量组成时,张量的非对角元素消失。本文第二部分对导致一对运动探测器输出的张量关系进行了空间积分。积分结果近似于一系列探测器对输出的总和。空间积分后的探测器张量将平移运动向量与合成输出向量联系起来。结果表明,运动向量与合成输出向量之间的夹角始终小于+/-90度,而运动向量与探测器对引发的局部响应向量之间的夹角可能覆盖整个角度范围。在本文的讨论部分,详细指出了运动计算场论的局限性及其更高阶近似。在一个特殊章节中讨论了探测器响应与模式属性的依赖关系,在另一个章节中讨论了与所谓孔径问题相关的问题。此外,在讨论中还提到了通过空间生理积分补偿模式相关偏差角的属性。