Proposal to reclassify , , and subsp. as comb. nov., comb. nov., comb. nov. and subsp. comb. nov., respectively and as a later heterotypic synonym of .

Recently, the members of the genus have been reclassified by Zheng At the same time, subsp. , , and have been successively and validly published, and has also been reclassified. In the present study, all these species and subspecies were re-evaluated by 16S rRNA gene sequence analysis or a combination of 16S rRNA gene sequence analysis and phylogenomic treeing. On the basis of the results presented here, we propose to reclassify , , and subsp. as comb. nov., comb. nov., comb. nov. and subsp. comb. nov., respectively and as a later heterotypic synonym of .