Four new species of subgenus from China.

subgenus is widely distributed in the world. In this study, 114 specimens were included in multigene phylogenetic analyses that allowed a better circumscription of the four sections in . subg. . Three new species from China, , and , are described and placed in different sections. The invalidly described is validated here as an additional new species. Comparisons between the sequences of the three closely related species , and support the distinction between these three species despite the paraphyly of , which results from its high intraspecific variability with numerous heteromorphisms.