Blickhan R
J Biomech. 1986;19(5):375-84. doi: 10.1016/0021-9290(86)90014-x.
The stiffness of the tibia-metatarsus joint of several spider-species was determined for different loading conditions. The relationship between force applied to the metatarsus tip and deflection is linear and free of hysteresis during axial loading (z'), but not under lateral (y') and dorsoventral (x') loading, where joint stiffness increases with amplitude and deflection rate. Up to lateral deflections of 2.5 degrees relaxation and nonlinearity of the joint-stiffness can be described by power functions. Considering strain induced in the spider's tibia, the viscoelastic properties of the joint result in a nonlinear amplitude transfer and a high-pass filtering of mechanical vibrations applied to the spider's leg. However, this contributes only slightly to the corresponding transfer characteristics measured for the electrical response of biological strain receptors.
针对几种蜘蛛物种，测定了胫跗关节在不同加载条件下的刚度。在轴向加载（z'）过程中，施加于跗尖的力与挠度之间的关系呈线性且无滞后现象，但在横向（y'）和背腹向（x'）加载时并非如此，此时关节刚度随振幅和挠度率增加。在高达2.5度的横向挠度下，关节刚度的松弛和非线性可用幂函数描述。考虑到蜘蛛胫节中产生的应变，关节的粘弹性特性导致了施加于蜘蛛腿部的机械振动的非线性振幅传递和高通滤波。然而，这对生物应变感受器电反应所测得的相应传递特性的贡献仅微乎其微。
