The extinct shark, (Elasmobranchii, Ptychodontidae) in the Upper Cretaceous of central-western Russia-The road to easternmost peri-Tethyan seas.

Isolated teeth belonging to the genus Agassiz, 1834 (Chondrichthyes; Elasmobranchii) from the Upper Cretaceous of the Ryazan and Moscow Oblast regions (European Russia) are described and discussed in detail herein. The taxonomic composition of the assemblage from the Ryazan region is very diverse including the first records of the cuspidate species . and . , which thus is largely consistent with those from other contemporaneous European localities. ubiquitously inhabited epicontinental seas of Europe during most of the Cretaceous with the most diverse assemblages coming from southern England, northern Italy, Belgium, and European Russia. Additionally, the material documented here from the Cenomanian of Varavinsky ravine area (Moscow Oblast) represents the northernmost occurrence of hitherto reported from Europe. It is evident that the Late Cretaceous shallow seas of the Russian platform represented a crucial pathway for the dispersal of from the European peri-Tethys to the eastern margins of the Neo-Tethyan Ocean. The Albian-Campanian records of from Europe indicate that its dominance in the peri-Tethys persisted for most of its evolutionary history. A local temperature drop across most of the European shallow seas probably contributed to the narrowing of its geographic range in the peri-Tethyan seas towards the end of the Mesozoic Era. The fossil remains of documented herein are accordingly of utmost importance for better understanding the taxonomic composition of Russian fossil ichthyofaunas and also inform about the dispersal of towards western and eastern peri-Tethyan seas during the Late Cretaceous.