McReynolds J S, Gorman A L
J Gen Physiol. 1970 Sep;56(3):376-91. doi: 10.1085/jgp.56.3.376.
Intracellular recordings were obtained from single visual cells of the scallop, Pecten irradians. Two types of units are found. One type gives a graded, depolarizing response to light and the other a graded, hyperpolarizing response. The depolarizing cells are 2-3 log units more sensitive to light and have a longer latency than the hyperpolarizing type. At high light intensities the depolarizing cells are inactivated while the hyperpolarizing cells maintain their responses. When action potentials are seen they occur during illumination in depolarizing cells ("on" response) and after illumination in hyperpolarizing cells ("off" response). The evidence suggests that the depolarizing responses are from the microvilli-brearing proximal cells, and the hyperpolarizing responses from the ciliary-type distal cells of the retina, and that both responses are directly produced by light.
从海湾扇贝（Pecten irradians）的单个视觉细胞中进行细胞内记录。发现了两种类型的细胞。一种类型对光产生分级的去极化反应，另一种产生分级的超极化反应。去极化细胞对光的敏感度比超极化细胞高2 - 3个对数单位，且潜伏期更长。在高光强度下，去极化细胞失活，而超极化细胞保持其反应。当出现动作电位时，它们在去极化细胞光照期间出现（“开”反应），在超极化细胞光照后出现（“关”反应）。证据表明，去极化反应来自视网膜中带有微绒毛的近端细胞，超极化反应来自睫状型远端细胞，并且两种反应均由光直接产生。
