Butler W F, Heap P
Histochem J. 1982 Jan;14(1):113-23. doi: 10.1007/BF01041134.
It has been confirmed that glycosaminoglycan is associated with collagen and other constituents of the anulus fibrosus. In cells, it appears to be present on the rough endoplasmic reticulum and is removed by hyaluronidase digestion and thus may consist of chondroitin sulphate. In pericellular material it appears as fibrils and granules, is unaffected by hyaluronidase and may consist of keratan sulphate or dermatan sulphate. Collagen fibrils contain hyaluronidase-digestable glycosaminoglycan (such as chondroitin sulphate) in the form of peripheral rings that appear as bands in longitudinal sections of the fibrils.
现已证实，糖胺聚糖与纤维环中的胶原蛋白及其他成分相关。在细胞中，它似乎存在于粗面内质网上，可被透明质酸酶消化去除，因此可能由硫酸软骨素组成。在细胞周围物质中，它以纤维和颗粒形式出现，不受透明质酸酶影响，可能由硫酸角质素或硫酸皮肤素组成。胶原纤维含有可被透明质酸酶消化的糖胺聚糖（如硫酸软骨素），呈外周环形式，在纤维的纵向切片中表现为条带。
