Department of Plant & Microbial Biology, University of California at Berkeley, 111 Koshland Hall, No. 3102, Berkeley, California 94720-3102.
Mycologia. 2003 May-Jun;95(3):442-56.
The position and composition of Macrolepiota within the Agaricaceae and its phylogenetic relationships with other members of the family were investigated, using both molecular (ITS and LSU rDNA sequences) and morphological characters. The molecular data separate the genus into two clades. The first clade comprises M. procera, M. mastoidea, M. clelandii and allies and is a sister group of Leucoagaricus and Leucocoprinus. The second, more diverse, clade, with M. rachodes and allies, M. globosa, Chlorophyllum molybdites, Leucoagaricus hortensis and Endoptychum agaricoides, is a sister group of Agaricus. The separation of the two clades is supported by morphological characters, such as the structure of the pileus covering, the stipitipellis and the shape of the germ pore and the spore apex. The two clades are regarded as genera for which the names Macrolepiota and Chlorophyllum are proposed. Macrolepiota nympharum does not belong to either clade but is assigned to the genus Leucoagaricus, close to L. leucothites. Endoptychum depressum is transferred to the genus Agaricus as A. inapertus.
使用分子（ITS 和 LSU rDNA 序列）和形态特征研究了 Macrolepiota 在伞菌科中的位置和组成及其与该科其他成员的系统发育关系。分子数据将该属分为两个分支。第一个分支包括 M. procera、M. mastoidea、M. clelandii 及其近缘种，是 Leucoagaricus 和 Leucocoprinus 的姊妹群。第二个分支更为多样化，包括 M. rachodes 及其近缘种、M. globosa、Chlorophyllum molybdites、Leucoagaricus hortensis 和 Endoptychum agaricoides，是 Agaricus 的姊妹群。两个分支的分离得到了形态特征的支持，例如菌盖覆盖物的结构、菌褶和担孢子顶端的形态。这两个分支被视为两个属，分别提出了 Macrolepiota 和 Chlorophyllum 的名称。Macrolepiota nympharum 不属于这两个分支，而是被归入 Leucoagaricus 属，与 L. leucothites 关系密切。Endoptychum depressum 被转移到 Agaricus 属作为 A. inapertus。
