Elven Hallvard, Bachmann Lutz, Gusarov Vladimir I
Hallvard Elven, National Center for Biosystematics, Natural History Museum, University of Oslo PO Box 1172 Blindern, NO-0318 Oslo, Norway.
Zool Scr. 2012 Nov;41(6):617-636. doi: 10.1111/j.1463-6409.2012.00553.x. Epub 2012 Jun 20.
Elven, E., Bachmann, L. & Gusarov V. I. (2012) Molecular phylogeny of the Athetini-Lomechusini-Ecitocharini clade of aleocharine rove beetles (Insecta). -Zoologica Scripta, 41, 617-636.It has previously been shown that the Aleocharinae tribes Athetini and Lomechusini form a well-supported clade, which also includes the small Neotropical tribe Ecitocharini. However, neither Athetini nor Lomechusini were recovered as monophyletic. In this study, we addressed the basal phylogenetic relationships among the three tribes using sequence data from (i) a mitochondrial fragment covering the COI, Leu2 and COII genes; (ii) a mitochondrial fragment covering part of the 16S gene, the Leu1 gene and part of the NADH 1 gene; and (iii) a part of the nuclear 18S gene, for 68 Athetini, 33 Lomechusini and 2 Ecitocharini species, plus representatives from 10 other tribes. The athetine subtribe Geostibina was recovered as sister group to the 'true Lomechusini', which included the type genus Lomechusa. The two clades formed a sister group to the main Athetini clade, which also included Ecitocharini and the 'false Lomechusini', a group of New World genera normally placed in Lomechusini. The following changes in classification are proposed: (i) Geostibina Seevers, 1978 is raised to tribal rank, and 13 Athetini genera are placed in Geostibini; (ii) Ecitodonia Seevers, 1965; Ecitopora Wasmann, 1887, and Tetradonia Wasmann, 1894 are moved from Lomechusini to Athetini; (iii) Ecitocharini Seevers, 1965 is placed in synonymy with Athetini; (iv) Discerota Mulsant & Rey, 1874 is tentatively included in Oxypodini; (v) Actocharina Bernhauer, 1907 is placed in synonymy with Hydrosmecta Thomson, 1858.
埃尔文，E.，巴赫曼，L. 及古萨罗夫，V. I.（2012年）隐翅虫亚科阿特西尼族 - 洛美切西尼族 - 埃西托查里尼族分支的分子系统发育研究（昆虫纲）。 - 《动物学文献》，41卷，617 - 636页。此前已有研究表明，隐翅虫亚科的阿特西尼族和洛美切西尼族形成了一个得到充分支持的分支，该分支还包括新热带区的小部落埃西托查里尼族。然而，阿特西尼族和洛美切西尼族都未被发现是单系的。在本研究中，我们利用以下序列数据探讨了这三个部落之间的基础系统发育关系：（i）一个覆盖细胞色素氧化酶亚基I（COI）、亮氨酸转运RNA2（Leu2）和细胞色素氧化酶亚基II（COII）基因的线粒体片段；（ii）一个覆盖16S基因部分、亮氨酸转运RNA1（Leu1）基因和烟酰胺腺嘌呤二核苷酸脱氢酶1（NADH 1）基因部分的线粒体片段；以及（iii）核18S基因的一部分，涉及68种阿特西尼族、33种洛美切西尼族和2种埃西托查里尼族物种，外加来自其他10个部落的代表物种。阿特西尼族的地锑亚族被发现是“真正的洛美切西尼族”的姐妹群，“真正的洛美切西尼族”包括模式属洛美丘萨属。这两个分支形成了主阿特西尼族分支的姐妹群，主阿特西尼族分支还包括埃西托查里尼族和“假洛美切西尼族”，“假洛美切西尼族”是一组通常被归入洛美切西尼族的新大陆属。现提出以下分类变动：（i）将西弗斯于1978年设立的地锑属提升为族级，13个阿特西尼族属被归入地锑族；（ii）将西弗斯于1965年设立的埃西托多尼亚属、瓦斯曼于1887年设立的埃西托波拉属和瓦斯曼于1894年设立的四齿属从洛美切西尼族移至阿特西尼族；（iii）将西弗斯于1965年设立的埃西托查里尼族归入阿特西尼族的同义词；（iv）暂时将穆尔桑特和雷伊于1874年设立的迪塞罗塔属归入尖足隐翅虫族；（v）将伯恩豪尔于1907年设立的阿克托查里纳属归入汤姆森于1858年设立的水湿隐翅虫属的同义词。
