Tamarin A, Lewis P, Askey J
J Morphol. 1976 Jun;149(2):199-221. doi: 10.1002/jmor.1051490205.
The byssus attachment plaque and the tissues responsible for its formation were studied in M. californianus by light microscopy and by transmission and scanning electron microscopy. It was shown that the plaque consists of at least three phases which ultrastructurally resemble three secretions considered to be collagen, mucoid material and polyphenol. The mucoid and polyphenol appear to mix as a colloidal suspension in which the latter is the continuous phase and forms the definitive bonding surface. Plaque collagen represents an extension of thread material into the cementing substance. Stimulated secretion within the ducts and distal depression of the mussel's foot shows a continuum of increasing heterogeneity from the inner toward the outer regions. This reflects the distribution of exocrine cell apices wherein exocytosis of polyphenol granules predominate deeply, mucous granules superficially and collagen granules in between. It is proposed that the morphology of the plaque conforms to theoretical physical-chemical requirements for adhesion under water.
利用光学显微镜、透射电子显微镜和扫描电子显微镜对加州贻贝的足丝附着板及其形成组织进行了研究。结果表明，附着板至少由三个阶段组成，其超微结构类似于被认为是胶原蛋白、黏液物质和多酚的三种分泌物。黏液物质和多酚似乎以胶体悬浮液的形式混合，其中多酚是连续相并形成最终的粘结表面。附着板胶原蛋白代表丝状物质向粘结物质的延伸。贻贝足部导管和远端凹陷内的刺激分泌显示，从内部到外部区域，异质性不断增加，呈现出连续变化。这反映了外分泌细胞顶端的分布情况，其中多酚颗粒的胞吐作用在深部占主导，黏液颗粒在表面占主导，胶原蛋白颗粒则介于两者之间。有人提出，附着板的形态符合水下粘附的理论物理化学要求。
