Phylogenomic analyses of the family suggest the reclassification of five species within the genus as heterotypic synonyms, the promotion of five subspecies to novel species, the taxonomic reassignment of five species to gen. nov., and the formal assignment of to the family .

Phylogenetic analyses based on 16S rRNA gene sequences of members of the family showed the presence of para- and polyphyletic genera. This finding prompted a thorough investigation into the taxonomy of the family by analysing their core genome phylogeny complemented with genome-based indices such as digital DNA-DNA hybridization, average nucleotide identity and average amino acid identity. The resulting data suggested the following proposals: was reduced in taxonomic rank as a later heterotypic synonym of ; subsp. as a later heterotypic synonym of subsp. ; the unification of subsp. and subsp. as ; the unification of subsp. and subsp. as ; the unification of subsp. and subsp. as ; subsp. as the novel species ; subsp. as the novel species ; subsp. as the novel species ; subsp. as the novel species ; subsp. as the novel species ; the reassignment of , , , and to the novel genus with as the type species; and the formal assignment of as a member of the family .